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Chamorro, William et al. Listado de especies y clave de géneros y subgéneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Rev. Colomb. Entomol., Jun 2018, vol.44, no.1, p.72-100. ISSN 0120-0488

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