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Biota colombiana

Print version ISSN 0124-5376On-line version ISSN 2539-200X

Biota colombiana vol.23 no.1 Bogotá Jan./June 2022  Epub Feb 09, 2022

https://doi.org/10.21068/2539200x.962 

Artículos

Diversity of Geadephaga (Coleoptera: Carabidae and Cicindelidae) in Colombia: an approach from existing literature

Diversidad de Geadephaga (Coleoptera: Carabidae y Cicindelidae) en Colombia: una aproximación desde la literatura existente

Anderson Arenas-Clavijo1 
http://orcid.org/0000-0001-5639-5273

James Montoya-Lerma2 
http://orcid.org/0000-0003-2122-1323

Pierre Moret3 
http://orcid.org/0000-0002-0141-4999

1Universidad del Valle Cali, Colombia ander-son.arenas@correounivalle.edu.co

2Universidad del Valle Cali, Colombiaja-mes.montoya@correounivalle.edu.co

3CNRS - Université Toulouse Jean Jaurès. Toulouse, France pierre.moret@univ-tlse2.fr

Abstract

The beetle families Carabidae and Cicindelidae, with about 40 000 species worldwide, play an increasing role in biodiversity conservation and ecological studies in Colombia, which makes a faunistic update all the more necessary. We present and analyze a checklist of the species so far described or registered from Colombia until 2020, based on published studies, and their distribution data at the department level. Carabidae are represented by 625 species and Cicindelidae by 100. Magdalena, Cundinamarca and Valle del Cauca are the departments with the highest record numbers of Carabidae, while Cicindelidae are most frequently cited from Valle del Cauca, Antioquia and Meta. The greater sampling effort in these regions is likely due to the accessibility of their natural areas from the main towns. The geographical distribution of Cicindelidae is better documented, with only 17 % of the species without mention of specific localities within the country, against 40 % in Carabidae. Hyper-diverse regions, such as the Chocó and the Amazon, are underrepresented. The number of species currently recorded is therefore estimated to be much lower than true diversity for both families.

Key words: Biodiversity; Faunistics; Ground beetles; Species list; Tiger beetles.

Resumen

Las familias Carabidae y Cicindelidae, con cerca de 40 000 especies a nivel mundial, tienen un papel creciente en estudios de biodiversidad y ecología en Colombia, lo que hace necesaria una puesta al día de los conocimientos sobre su diversidad. Presentamos y analizamos la lista de las especies descritas/registradas en Colombia hasta 2020, con base en estudios publicados, y sus datos de distribución a escala de departamento. Los Carabidae están representados por 625 especies y los Cicindelidae por 100. Magdalena, Cundinamarca y Valle del Cauca presentan los mayores números de registros de Carabidae, mientras que para Cicindelidae son Valle del Cauca, Antioquia y Meta. Probablemente, el mayor esfuerzo de muestreo en estas regiones se debe a la accesibilidad de sus áreas naturales desde las principales ciudades. Los Cicindelidae se benefician de más registros de localidad, con solo 17 % de las especies sin mención de localidad específica en el territorio nacional, contra el 40 % en Carabidae. Regiones hiperdiversas, como el Chocó y la Amazonía, están subrepresentadas. Por consiguiente, se estima que el número de especies actualmente citadas es muy inferior a la diversidad real de las dos familias.

Palabras clave: Biodiversidad; Escarabajos del suelo; Escarabajos tigre; Faunística; Lista de especies.

Introduction

Geadephaga, the group of terrestrial Adephaga formed by Carabidae (ground beetles), Cicindelidae (tiger beetles) and Trachypachidae, with about 40 000 described species worldwide, represents nearly 10 % of the species so far described in the order Coleoptera (Bouchard et al., 2017; Lorenz, 2017). Within this monophyletic group (López-López & Vogler, 2017), the position of tiger beetles is still under debate, whether as Cicindelidae at family rank (López-López & Vogler, 2017; Duran & Gough, 2020) or as Cicindelinae within Carabidae (Gough et al., 2019; Li et al., 2020). In this study, we provisionally placed tiger beetles at family rank without prejudice to future advances in genomic studies, because in most studies they have been treated separately from ground beetles. Trachypachidae, on their part, are not represented in the Neotropics.

Carabidae and Cicindelidae are present in almost every possible ecosystem worldwide (Lövei & Sunderland, 1996), with a great diversity of life history traits, and frequently used as bioindicators in ecological studies (Rainio & Niemelä, 2003; Pearce & Venier, 2006) or as control agents against agricultural pests, due to their predatory habits (Kromp, 1999; de Heij & Willenborg, 2020). Therefore, their knowledge can be crucial in studies on sustainable agriculture, or on assessment of vulnerable natural areas.

Information on Geadephaga of Colombia has been synthetized in two studies more than fifteen years ago: one by Vítolo (2004) on Cicindelidae, and another by Martínez (2005) on Carabidae. Vítolo (2004) offers data on morphology, distribution in Colombia, identification keys, and a summary of the diversity of tiger beetles in the country. Martínez (2005) provides identification keys with full habitus illustrations for all the genera present in Colombia. As the main objective of the latter study was to allow identification at the genus level, the information on the distribution of taxa was also limited to genera. Moreover, the list of species in Martínez (2005) needs revision in three aspects: some species cited from Colombia in the scientific literature were not included, the presence in Colombia of other species is stated without references, and a few unnamed morphospecies are listed, which hinders a correct evaluation of the total number of species cited from Colombia. As a complement to these fundamental works, Moret (2003) published an identification key to the Carabidae of the Andean paramo which includes southern Colombia, but it is also limited to genera and does not provide any distributional information.

In Colombia, the uncertainty about the identity of many species (Linnaean shortfall) and their spatial distribution (Wallacean shortfall) are the greatest obstacles to develop robust ecological studies (Hortal et al., 2015). In addition, poor taxonomic and faunistic knowledge limits the potential assessment of the ecosystem services that these beetles could provide. Parataxonomy (i.e. the assignation of individuals to morphospecies based on their external appearance, without considering neither taxonomic literature nor the specialists within the group), as an alternative way to overcome the taxonomic impediment in ecological studies, appears to be an unsatisfactory solution, since it tends to overestimate the number of species and does not guarantee reproducibility (Krell, 2004).

The knowledge of Colombian Geadephaga has increased in recent years, due to a growing number of taxonomic (e.g. Arenas-Clavijo, 2017; Forero et al., 2019; Moret, 2019; Sarmiento-Roa et al., 2020) and ecological studies (e.g. Arenas & Armbrecht, 2019; García-Suabita et al., 2019) based on this group during the last decade. An update of the current taxonomic and faunistic knowledge was therefore necessary, as a starting point for future systematic revisions of the tribes or genera which might deserve special attention from a biogeographical or ecological point of view.

For these reasons, the present study aims to list the Carabidae and Cicindelidae species recorded from Colombia in the literature, up to 2020, thus offering a current state of knowledge of these two families, emphasizing gaps and biases. It also seeks to provide information on areas of potential interest, where it would be convenient to intensify, complete or initiate samplings of these ecologically important beetles.

Material and methods

Bibliographic database

A total of 150 studies published up to 2020, related to ground and tiger beetles of Colombia, was obtained from an exhaustive search that included documents from the 19th century. Much of this literature is available in electronic repositories (e.g. Biodiversity Heritage Library, Gallica, Persée), and a smaller portion was gathered by direct request to libraries or authors. The reviewed publications cover the following topics: (1) descriptions of new species based on material from Colombia or, in the mid-19th century studies, from “Nueva Granada” or “Nouvelle Grenade”; (2) new Colombian records of species already described from other countries; (3) lists of species of a genus or of a supra-generic taxon in systematic revisions; and (4) ecological studies or biodiversity inventories. Regarding the last category, doubtful attributions marked as “cf.” or “aff.”, as well as morphospecies designated by a number, were not included. Based on this inventory, the references of all the species listed by Martínez (2005) were verified (Appendix 1), and a new checklist was produced.

Construction of the checklist

Once the references were compiled, a list was constructed in which species are sorted by genus. Infraspecific taxa were excluded from the list. Generic and subgeneric names were updated in accordance with Lorenz (2017) or based on the last available systematic revisions. In the case of synonyms or debated generic attribution, we followed as a general rule the taxonomic decision of the most recent revision, the reference of which is given in the bibliographic section of the list (Appendix 2, last column). Locality information is limited to the “Departamento” (department or province), and only published localities were taken into account.

Data analysis

The checklist was used as a database to build graphs and maps to better understand the current state of knowledge of the group in Colombia. These graphs are based on the description year or on the year of the first record for each species, on the number of species per genus, and on the number of species per department.

Results

Carabidae

Based on published data, the family Carabidae is currently represented in Colombia by 625 species -close to 1.7 % of the world total-, arranged in 10 subfamilies, 32 tribes and 125 genera (Appendix 2). Of the recorded species, 330 (53 %) have not been reported from any other country, so that could be considered Colombian endemics, at least until they do not get recorded from any surrounding country. Thirteen genera account for 50 % of the species, the most speciose genus being DyscolusDejean, 1831 (Harpalinae, Platynini), which contains 14 % of the total (85 species) (Figure 1). Forty-seven genera (38 %) are represented by a single species in Colombia (Table 1), five of which are monospecific, with records of their unique species in at least another country: Stenocheila (Harpalinae, Lachnophorini); Askalaphium (Harpalinae, Ctenodactylini); Enceladus (Siagoninae, Siagonini); Homalomorpha (Harpalinae, Cratocerini) and Trichognatha (Harpalinae, Galeritini) (Martínez 2003, 2005; Martínez & Ball, 2003; Lorenz, 2017). Three genera are only known from Colombia: Callidadelpha (Harpalinae, Lebiini), Cryptomma (Scaritinae, Clivinini) (without specific locality recorded) and Columbitrechus (the latter, most likely a synonym of Paratrechus). Finally, two introduced species have been registered: Laemostenus complanatus, from northern Africa and/or western Europe (Martínez, 2005), and Mochtherus tetraspilotus, from southeastern Asia (Torres-Domínguez et al., 2020).

Figure 1 Number and proportion of species by genus of ground beetles recorded from Colombia 

Table 1 Structure of the subfamilies and tribes of Carabidae Latreille, 1802 in Colombia. In parentheses: number of genera with only one species recorded in the country. 

Regarding species geographic distribution, 254 species (~41 %) are known from the country as a whole without more precise localities, from which 143 have not been cited from any other country; 371 species (~59 %) have at least one specific locality record. 309 species are known from a single department (Figure 4), and 163 of these are exclusive to Colombia. The departments with the highest number of recorded species are: Magdalena (79), Cundinamarca (75) and Valle del Cauca (43), while Guaviare, San Andrés y Providencia and Sucre have no records for the family (Appendix 1, Figures 2 and 3).

Figure 2 Number of ground beetle species per Colombian departments. 

Figure 3 Number of ground beetle species per Colombian department, based on data published from 1831 to 2020. 

Figure 4 Number and proportion of species by genus of tiger beetle recorded from Colombia. 

The species with the widest distribution in Colombia are Laemostenus (Laemostenus) complanatus (Harpalinae, Sphodrini) and Pelmatellus variipes (Harpalinae, Harpalini), each one with locality data in 11 departments (Camero, 2003; Martínez & Ball, 2003), but most of the species with the widest distribution belong to the tribe Platynini. However, in some cases, systematic revisions could reveal the existence of several species with more limited distributions under the currently accepted name.

Cicindelidae

The family Cicindelidae is currently represented in Colombia by 100 species distributed in three tribes and 21 genera of the subfamily Cicindelinae (Appendix 3). This figure represents 3.3 % of the world total of tiger beetle species, a percentage twice as high as in Carabidae. Of the species registered in the Colombian territory, 16 do not have records in any other country, most of them probably endemic according to the available data. The most diverse genus is Odontocheila, with 16 species, while 81 % of the remaining species is contained in seven different genera (Figure 4).

Eight tiger beetle genera are represented by a single species in Colombia (Table 2), but none has a distribution limited to the country. Callidema boussingaulti (Cicindelini) has a restricted distribution in mountain environments, since it is known only from the Andes of Colombia, Peru and Ecuador (Cassola & Pearson, 2001). Other genera such as Cheiloxya (Cicindelini), Cenothyla (Cicindelini) and Ronhuberia (Cicindelini) are distributed in northern South America (Cassola & Pearson, 2001; Moravec, 2020).

Table 2 Structure of the subfamilies and tribes of Cicindelidae Latreille, 1802 in Colombia. In parentheses: number of genera with only one species recorded in the country. 

Regarding the distribution of Cicindelidae in Colombia, 17 species (17 %) do not have any precise locality records, three of which appear to be endemic to the country: Odontocheila simulator, although the holotype and only known specimen of this dubious taxon is probably an artefact (Moravec, 2018); Pseudoxycheila ceratoma; and Pseudoxycheila tarsalis (Cicindelini). 21 species are known from only one department (Figure 5), and of these, four are not recorded from other countries: Odontocheila hamulipenis and Oxycheila pseudoaquatica (Cicindelini) from Valle del Cauca; Oxygonia kippenhani from Boyacá; and Ctenostoma (Procephalus) maculosum (Ctenostomatini) from Cundinamarca (Appendix 3).

Figure 5 Distribution of tiger beetle species in Colombia, per departments. 

The departments with the highest number of tiger beetle species records are Valle del Cauca (36), Antioquia (27) and Meta (27), whereas San Andrés, Providencia and Arauca do not have records (Appendix 3 and Figure 6). Vaupés department, with only one record (Odontocheila trilbyana), is located in the Amazonia, a region where many tiger beetle species have already been reported in neighboring countries, which highlights the low level of sampling of this ecosystem in Colombia.

Figure 6 Number of tiger beetle species per Colombian department, based on data published from 1842 to 2020. 

The species with the widest distribution in Colombia are Pseudoxycheila bipustulata, with records in 20 departments, and Tetracha (Tetracha) sobrina in 19 (Appendix 3). Regarding the genus PseudoxycheilaVítolo (2004) expressed doubts about the status of some species proposed by Cassola (1997), as species differentiation is especially difficult in this Andean genus, and several of the new species described by Cassola appear to fall within the morphological variation range of P. bipustulata. In this case, as in many others, a molecular approach would be necessary to solve the issues left by conventional morphology-based taxonomy.

Discussion

The current knowledge of the families Carabidae and Cicindelidae in Colombia is the result of a two centuries-long history (Figure 7).Regarding Carabidae, taxonomic descriptions based on material from Colombia began as early as 1825,but they reached their greatest development between the second third and the end of the 19th century, a period in which 70 % of the carabid fauna known from the country had already been described, mainly thanks to the contributions of Louis Reiche (52 species described- between 1842 and 1843), Maximilien de Chaudoir (131 species described between 1848 and 1880) and Jules Putzeys (89 species described between 1846 and 1878). In this period, most descriptions were very brief, not illustrated, with limited distribution data or without any, and they did not fit the current standards of insect taxonomy. Furthermore, very few have been subject to taxonomic revisions in recent times, which makes their identification difficult, if not impossible without examining type specimens.

Figure 7 Cumulative number of ground beetle species described (in blue) and recorded from Colombia (in orange), in five-years increments, up to 2020. Species described from Colombia but subsequently synonymized are not included. 

The genera in which this situation is especially detrimental are Dyscolus, Notiobia, Lebia and Agra,among others. From the beginning of the twentieth century, the rate of description of new species has been very slow, with a slight rebound from 1980. The new records from Colombia (i.e. for species whose description was based on specimens from other countries) have followed a much slower pace, with a gradual increase from the mid-20th century, when authors such as Stefano Straneo (records from Colombia from 1951 to 1991), Joaquim Mateu (1961-1998), George E. Ball (1978-2013), Terry L. Erwin (1970-2016), Danny Shpeley (1978-2013) , among others, contributed to the knowledge of the fauna of Colombia.

The investigations on Cicindelidae followed a different path (Figure 8). The pace of descriptions was globally slow, with two acceleration points: a weak one during the second third of the 19th century, with contributions by several authors who published isolated species descriptions, and another very strong in the last decade of the twentieth century, based on more comprehensive works, especially revisions of genera at a continent scale, such as those by Fabio Cassola (1997), Roger Naviaux (1998), Jürgen Wiesner (1992), 1999). Thanks to these taxonomic revisions and to several local, national or continental checklists (Cassola & Pearson, 2001; Fernández et al., 1993; Moravec 2018, 2020; Vítolo & Pearson, 2003; Wiesner, 2020), Colombian records of species described from other countries increased dramatically since 1990, exceeding the number of new descriptions. In this regard, tiger beetle knowledge has progressed much faster in recent decades than that of ground beetles. This greater sampling and publication effort may also explain why the Colombian records amount to 3.3 % of the world total of tiger beetle species, a percentage twice as high as in Carabidae.

Figure 8 Cumulative number of tiger beetle species described from Colombia (in blue) and recorded from Colombia (in orange), in five-year increments, up to 2020. Species described from Colombia but subsequently synonymized are not included. 

The geographical distribution of the locality records of Carabidae (Figure 3) is largely due to accessibility factors. In most cases, naturalists of the 19th century and the first half of the 20th century were constrained by the reduced transportation network of that time and collected along the same routes around the main cities. Natural areas near main roads and navigable rivers were the most surveyed, which creates an over-sampling bias and makes it difficult to assess the true diversity of each region, in line with the results of a study performed in a neighboring country (Donoso et al., 2009). Another kind of bias derives from the activity of a particular researcher in a limited area. For example, most of the species reported from the Magdalena department correspond to species of the genus Blennidus, described by Stefano Straneo from specimens collected by Philip J. Darlington in the Sierra Nevada de Santa Marta in 1928 and 1929.Conversely, ground beetle diversity is clearly underestimated in less accessible hyper-diverse regions such as the Chocó ecoregion, the Amazon foothills (Caquetá and Putumayo departments) and the Amazon rainforest itself (Guainía, Guaviare and Vaupés departments). Although true diversity cannot be precisely assessed based on available data, it is expected that these regions have a more diverse carabid fauna than what is currently known.

The distribution of tiger beetle records throughout the Colombian territory (Figure 6) is more balanced than that of ground beetles, possibly due to the interest the group has provoked among professional and amateur entomologists over the last decades.

When comparing the number of Geadephaga species recorded so far from Colombia (725) with that of other countries, it appears clearly that there is still a huge work ahead before approaching a complete knowledge of their diversity in this country.

Our results in Colombia are comparable to those obtained in Peru, where 690 species are currently known (Erwin et al., 2015, leaving aside unnamed morphospecies only designated by numbers) in a territory of a size similar to that of Colombia. In Brazil, a territory eight times larger, 1506 species have been reported, that is, only twice as many (Roig Juñent & Domínguez, 2001).

The level of knowledge is higher in Ecuador, with 763 species (Moret & Salazar, unpublished data) in a territory five times smaller and in ecosystems shared with southern Colombia: Chocó ecoregion, Andean montane forest, paramo, interandean dry shrublands or woodlands, Amazonian lowlands. Outside the Neotropics, to mention just one example, 693 species have been reported in the state of Texas in the United States of America (Bousquet, 2012), a territory twice as small.

The magnitude of the work to be done is obvious when noting that the late Terry L. Erwin collected more than 600 species of Carabidae (most of which remain undescribed) in the Pakitza reserve of Peruvian Amazonia, in a space of only 40 km2 (Erwin, 1991), that is, a number close to the total of the described species registered for the whole of Peru. In Colombia, the number of recorded species does not exceed 42 in any of the six departments (Nariño, Putumayo, Caquetá, Cauca, Huila, Meta) which contain environments similar to that of Pakitza, i.e. lower montane forest at an elevation of about 650 m in the Amazonian foothills of the Andes.

Conclusion and perspectives

The results of this study suggest that the deficient taxonomic and chorological knowledge of the families Carabidae and Cicindelidae in Colombia will be an obstacle to developing well supported ecological studies of this group, as long as species distributions are not accurately known. Publication of distributional information based on reliable identifications, as well as taxonomic revisions of the genera offering the highest value as bioindicators or pest control, are therefore urgent priorities.

Another challenge faced by the study of Geadephaga in Colombia is the possibility of reaching the specific level in identifications, an issue that can be of great importance in ecological studies. The identification keys currently available for Colombia (Moret, 2003; Martínez, 2005; Vítolo, 2004) are limited to the genus level. Species-level keys exist for a number of neotropical genera (e.g. Arndt, 1998; Ball & Shpeley, 2009; Boyd & Erwin, 2016), but they include species not found in Colombia and are therefore difficult to use by non-taxonomic biologists. Developing keys for all genera of ground beetles and tiger beetles recorded in Colombia is currently an unachievable goal, but it would be desirable to focus efforts on a small number of genera including a sufficient number of described species and of interest as bioindicators.

Finally, in the light of current knowledge and the available techniques, two paths are needed to improve future studies of ground beetles in Colombia: (1) compare specimens deposited in Colombian collections with types kept in European and North American museums, to generate valid vouchers for local taxonomists, and (2) retrieve molecular data (e.g. COI-based DNA barcodes) from these local vouchers, to facilitate future research.

Acknowledgments

We warmly thank Jiri Moravec for providing recent literature on Odontocheilina (Cicindelidae), Petr Bulirsch for useful comments on the systematics of Scaritinae (Carabidae), and Fernanda Salazar for her help in the construction of the database and information on the Ecuadorian carabid fauna. We especially thank Dione Seriperri, Museu de Zoologia, Universidade de São Paulo, for her help in finding publications by Hans Reichardt. AAC’s doctoral studies were partly funded by the “Fondo nacional de financiamiento para la ciencia, la tecnología y la innovación, Francisco José de Caldas”, contract number 491-2020 in program 1106-852-70306 “Relaciones multiescalares de la diversidad en gradientes altitudinales del bosque tropical”, project 70899 “Diversidad de artrópodos en gradientes altitudinales: una aproximación integrativa morfológica y molecular”. Finally, but not last, we thank the comments of two anonymous reviewers, which helped to improve the manuscript.

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Citación del artículo: Arenas-Clavijo, A., Montoya-Lerma, J. & Moret, P. (2022). Diversity of Geadephaga (Coleoptera: Carabidae and Cicin-delidae) in Colombia: an approach from existing literature. Biota Co-lombiana, 22(2), e962

Appendix 1. Bibliographic references reviewed to generate the species list of Geadephaga present in Colombia.

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Arenas-Clavijo, A. (2017). Primer registro de Notiobia (Notiobia) umbrifera Bates y redescubrimiento de Notiobia (Anisotarsus) praeclara Putzeys (Coleoptera: Carabidae) en Colombia. Boletín Científico del Centro de Museos. Museo de Historia Natural, 21(1), 226-233.

Arenas-Clavijo, A. (2018). Escarabajos tigre (Coleoptera: Cicindelidae) del Museo de Entomología de la Universidad del Valle, Colombia. Boletín del Museo de Entomología de la Universidad del Valle, 18(1), 32-45.

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Appendix 2 List of species of Carabidae Latreille, 1802 recorded from Colombia until 2020, with their respective distribution within the country. Species marked with asterisk (*) have been cited only from Colombia. Departments are codified according to ISO rule 3166-2. References are listed in appendix 1  

Subfamily Tribe Genus Species Department Reference
Brachininae Bonelli, 1810 Brachininae Bonelli, 1810 Brachininae Weber, 1801 Brachinus (Neobrachinus) aeger Chaudoir, 1876 No data Erwin, 1970
Brachinus (Neobrachinus) geniculatus Dejean, 1831 BOL Dejean, 1831
Brachinus (Neobrachinus) lateralis Dejean, 1831 No data Bates, 1883
Brachinus (Neobrachinus) limbiger Chaudoir, 1876 No data Chaudoir, 1876a
Brachinus (Neobrachinus) olidus Reiche, 1843* No data Reiche, 1843a
Pheropsophus Solier, 1833 Pheropsophus (Pheropsophus) aequinoctialis (Linnaeus, 1763) MAG Bates, 1883; Arenas-Clavijo & Posso-Gómez, 2017
Pheropsophus (Pheropsophus) rivierii (Demay, 1838) No data Chaudoir, 1876a
Carabinae Latreille, 1802 Carabini Latreille, 1802 Calosoma Weber, 1801 Calosoma (Carabosoma) angulatum Chevrolat, 1834 MAG Gidaspow, 1963
Calosoma (Carabosoma) glabratum Dejean, 1831 CUN Gidaspow, 1963
Calosoma (Castrida) abbreviatum Chaudoir, 1869 VAC Gidaspow, 1963
Calosoma (Castrida) alternans (Fabricius, 1792) VAC Gidaspow, 1963; Arenas-Clavijo & Posso-Gómez, 2017
Calosoma (Castrida) fulgens Chaudoir, 1869 VAC Gidaspow, 1963
Harpalinae Bonelli, 1810 Cratocerini Lacordaire, 1854 Catapiesis Solier, 1835 Catapiesis attenuata (Chaudoir, 1862) No data Reichardt, 1973
Catapiesis brasiliensis (Gray, 1832) No data Reichardt, 1973
Catapiesis columbica Chevrolat, 1838* No data Reichardt, 1973
Catapiesis tumida Reichardt, 1973* CUN Reichardt, 1973
Cratocerus Dejean, 1829 Cratocerus indupalmensis Grzymala & Will, 2014 CES Grzymala & Will, 2014
Homalomorpha Brullé, 1835 Homalomorpha castanea Brullé, 1835 No data Chaudoir, 1852
Chaetogenyini Bonelli, 1813 Camptotoma Reiche, 1843 Camptotoma lebasii Reiche, 1843* No data Reiche, 1843b
Harpalinae Bonelli, 1810 Ctenodactylini Laporte de Castelnau, 1834 Amblycoleus Chaudoir, 1872 Amblycoleus platyderus (Chaudoir, 1861) COR Chaudoir, 1861
Askalaphium Liebke, 1938 Askalaphium depressum (Bates, 1871) PUT Martínez, 2003
Leptotrachelus Latreille, 1829 Leptotrachelus aequinoctialis (Chaudoir, 1848)* No data Chaudoir, 1848
Leptotrachelus fulvicollis Reiche, 1842* No data Reiche, 1842b
Leptotrachelus setulosus Liebke, 1928 No data Mroczkowski, 1960
Leptotrachelus testaceus Dejean, 1831 BOL Dejean, 1831; Chaudoir, 1848
Cyclosomini Laporte de Castelnau, 1834 Anaulacus MacLeay, 1825 Anaulacus (Aephnidius) piceolus (Chaudoir, 1876) BOL, MAG Ball & Shpeley, 2002
Anaulacus (Macracanthus) ciliatus (Mutchler, 1934) VID Ball & Shpeley, 2002
Anaulacus (Macracanthus) sericatus (Chaudoir, 1846) MAG Ball & Shpeley, 2002
Tetragonoderus Dejean, 1829 Tetragonoderus (Peronoscelis) undatus Dejean, 1829 CAQ Martínez, 2003
Tetragonoderus (Peronoscelis) variegatus Dejean, 1829 No data Chaudoir, 1876b
Tetragonoderus (Peronoscelis) velutinus Motschulsky, 1864 GUA Martínez, 2003
Tetragonoderus (Tetragonoderus) quadriguttatus Dejean, 1829 HUI, MAG, TOL Martínez, 2003; Shpeley & Ball, 2008
Tetragonoderus (Tetragonoderus) tesselatus Chaudoir, 1876 MAG Martínez, 2003
Dercylini Bonelli, 1813 Dercylus Laporte de Castelnau, 1832 Dercylus (Dercylus) alternans Kuntzen, 1912 No data Kuntzen, 1912
Dercylus (Dercylus) opacus Kuntzen, 1912 No data Kuntzen, 1912
Dercylus (Licinodercylus) tuberculatus (Chaudoir, 1883)* CAL, CAU Moret & Bousquet, 1995
Galeritini LeConte, 1853 Galerita Fabricius, 1801 Galerita (Galerita) aequinoctialis Chaudoir, 1852 BOL Martínez, 2003
Galerita (Galerita) amazonica Liebke, 1939 ARA, BOY, CAS, CUN, MAG, MET, VID Camero, 2003
Galerita (Galerita) americana (Linnaeus, 1758) VAC, VID Arenas et al., 2013, García-Suabita et al., 2019
Harpalinae Bonelli, 1810 Galerita Fabricius, 1801 Galerita (Galerita) moritzi Mannerheim, 1837 MET, HUI, VAC Reichardt, 1967; Arenas-Clavijo & Posso-Gómez, 2017
Galerita (Galerita) mustelina Bates, 1884 RIS Reichardt, 1967
Galerita (Galerita) occidentalis (Olivier, 1795) RIS, VAC Reichardt, 1967; Arenas-Clavijo & Posso-Gómez, 2017
Galerita (Galerita) tristis Reiche, 1842 ATL, MAG Reichardt, 1967
Trichognatha Latreille, 1829 Trichognatha marginipennis Latreille, 1829 BOY Reichardt, 1967; Steinheil, 1875a
Harpalini Bonelli, 1810 Amblygnathus Dejean, 1829 Amblygnathus darlingtoni Ball & Maddison, 1987 MAG Ball & Maddison, 1987
Amblygnathus janthinus Dejean, 1829 No data Ball & Maddison, 1987
Amblygnathus suturalis Putzeys, 1845 ATL Sarmiento-Roa et al., 2020
Anisocnemus Chaudoir, 1843 Anisocnemus amblygonus Shpeley & Ball, 1978 ATL, CES Shpeley & Ball, 1978
Anisocnemus validus Chaudoir, 1843* No data Shpeley & Ball, 1978
Athrostictus Bates, 1878 Athrostictus circumfusus (Putzeys, 1878)* No data Putzeys, 1878a
Athrostictus luridus (Reiche, 1843) No data Putzeys, 1878a
Athrostictus paganus (Dejean, 1831) BOL, CAU Dejean, 1831; Putzeys, 1878a; Arenas & Armbrecht, 2019
Athrostictus velutinus (Putzeys, 1878)* No data Putzeys, 1878a
Bradycellus Erichson, 1837 Bradycellus (Bradycellus) apicalis Putzeys, 1878* CUN Putzeys, 1878b
Bradycellus (Bradycellus) celeripes Putzeys, 1878* CUN Putzeys, 1878b
Bradycellus (Bradycellus) suturiger Putzeys, 1878* ATL Putzeys, 1878b
Neoaulacoryssus Noonan, 1985 Neoaulacoryssus cupripennis (Gory, 1833) TOL Sarmiento-Roa et al., 2020
Notiobia Perty, 1830 Notiobia (Anisotarsus) praeclara Putzeys, 1878* CAU, CUN, VAC Noonan, 1981; Arenas-Clavijo, 2017
Notiobia (Notiobia) aeneola Putzeys, 1878* ATL Putzeys, 1878b
Harpalinae Bonelli, 1810 Harpalini Bonelli, 1810 Notiobia Perty, 1830 Notiobia (Notiobia) aulica (Dejean, 1829) ARA, BOY, CAU, CES, CUN, MAG, QUI, TOL Arndt, 1998; Camero, 2003
Notiobia (Notiobia) concolor Putzeys, 1878* CAL, CUN, NSA Putzeys, 1878b
Notiobia (Notiobia) disparilis Bates, 1878 CAQ Sarmiento-Roa et al., 2020
Notiobia (Notiobia) dubia Putzeys, 1878* ATL Putzeys, 1878b
Notiobia (Notiobia) glabrata Arndt, 1998 CAQ Sarmiento-Roa et al., 2020
Notiobia (Notiobia) jucunda Putzeys, 1878 CUN Putzeys, 1878b
Notiobia (Notiobia) longipennis Putzeys, 1878* No data Putzeys, 1878b
Notiobia (Notiobia) similis Putzeys, 1878* CAL, CUN Putzeys, 1878b
Notiobia (Notiobia) umbrifera Bates, 1884 CAU, VAC Arenas-Clavijo, 2017
Pelmatellus Bates, 1882 Pelmatellus columbianus (Reiche, 1843) BOY, CAL, CUN, SAN Moret, 2000
Pelmatellus variipes Bates, 1891 AMA, ARA, BOY, CAL, CUN, MAG, MET, QUI, SAN, VAU, VID Camero, 2003
Polpochila Solier, 1849 Polpochila (Polpochila) marginalis Nègre, 1963 MAG Nègre, 1963
Selenophorus Dejean, 1829 Selenophorus (Celiamorphus) discopunctatus Dejean, 1829 No data Reiche, 1843c
Selenophorus (Selenophorus) aurichalceus Dejean, 1831* BOL Dejean, 1831; Putzeys, 1878a
Selenophorus (Selenophorus) coracinus Dejean, 1831* BOL Dejean, 1831
Selenophorus (Selenophorus) cyaneus Putzeys, 1878* No data Putzeys, 1878a
Selenophorus (Selenophorus) dilutipes Putzeys, 1878 No data Putzeys, 1878a
Selenophorus (Selenophorus) dimidiatulus van Emden, 1958* MAG van Emden, 1958
Selenophorus (Selenophorus) distinctus Putzeys, 1878* No data Putzeys, 1878a
Selenophorus (Selenophorus) exilis Dejean, 1831* BOL, MAG Dejean, 1831; Putzeys, 1878a
Harpalinae Bonelli, 1810 Harpalini Bonelli, 1810 Selenophorus Dejean, 1829 Selenophorus (Selenophorus) irideus Reiche, 1843* No data Reiche, 1843c
Selenophorus (Selenophorus) irinus (Reiche, 1843) No data Reiche, 1843d; Putzeys, 1878a
Selenophorus (Selenophorus) laevicollis (Bates, 1884) No data Bates, 1883
Selenophorus (Selenophorus) liodiscus Putzeys, 1878* No data Putzeys, 1878a
Selenophorus (Selenophorus) modestus Putzeys, 1878* No data Putzeys, 1878a
Selenophorus (Selenophorus) myrmidon Dejean, 1831* BOL Dejean, 1831; Putzeys, 1878a
Selenophorus (Selenophorus) pyritosus Dejean, 1829 MAG, BOL Martínez, 2003
Selenophorus (Selenophorus) ruficollis (Putzeys, 1878) AMA Ball & Maddison, 1987
Selenophorus (Selenophorus) splendidus Putzeys, 1878 No data Putzeys, 1878a
Selenophorus (Selenophorus) striatopunctatus Putzeys, 1878 BOL Martínez, 2003
Selenophorus (Selenophorus) subaeneus Reiche, 1843 No data Reiche, 1843c
Selenophorus (Selenophorus) vicinus Dejean, 1829 No data Putzeys, 1878a
Selenophorus (Selenophorus) xantholomus Putzeys, 1878 No data Putzeys, 1878a
Selenophorus (Selenophorus) affinis Dejean, 1831 No data Putzeys, 1878a
Selenophorus (Selenophorus) brevis (Putzeys, 1878)* TOL Putzeys, 1878a
Stenomorphus Dejean, 1831 Stenomorphus angustatus Dejean, 1831 BOL Ball et al., 1991
Trichopselaphus Chaudoir, 1843 Trichopselaphus magnificus Ball, 1978 VAC Sarmiento-Roa et al., 2020
Helluonini Bonelli, 1813 Dailodontus Reiche, 1843 Dailodontus clandestinus (Klug, 1834) MAG, VAC Reichardt, 1974b; Arenas-Clavijo & Chacón de Ulloa, 2016
Helluobrochus Reichardt, 1974 Helluobrochus cribratus (Reiche, 1843) AMA, BOY, VID Reiche, 1843a; Reichardt, 1974b
Helluobrochus subrostratus (Bates, 1871) CAS Martínez, 2003
Helluomorphoides Ball, 1951 Helluomorphoides glabratus (Bates, 1871) No data Reichardt, 1974b
Helluomorphoides unicolor (Brullé, 1838) AMA Martínez, 2003
Harpalinae Bonelli, 1810 Helluonini Bonelli, 1813 Pleuracanthus Gray, 1832 Pleuracanthus sulcipennis Gray, 1832 MET Martínez, 2003
Lachnophorini LeConte, 1853 Amphithasus Bates, 1871 Amphithasus elegans (Dejean, 1831)* BOL Zamorano et al., 2019
Anchonoderus Reiche, 1843 Anchonoderus apicalis Reiche, 1843* No data Reiche, 1843b
Anchonoderus binotatus Reiche, 1843 No data Reiche, 1843b
Anchonoderus cyanescens (Putzeys, 1878)* BOY Putzeys, 1878b; Zamorano et al., 2019
Anchonoderus erosus Putzeys, 1878* No data Putzeys, 1878b
Anchonoderus femoratus Putzeys, 1878* BOY Putzeys, 1878b
Anchonoderus myops Reiche, 1843 No data Reiche, 1843b
Anchonoderus reichei Putzeys, 1878* ANT, BOY, CUN NSA Putzeys, 1878b
Anchonoderus subaeneus Reiche, 1843 No data Reiche, 1843b
Anchonoderus unicolor Chaudoir, 1850* No data Chaudoir, 1850
Asklepia Liebke, 1938 Asklepia laetitia Zamorano & Erwin, 2014* AMA Erwin & Zamorano, 2014
Calybe Laporte de Castelnau, 1834 Calybe grata (Motschulsky, 1864) No data Motschulsky, 1864
Ega Laporte de Castelnau, 1835 Ega aequatoria Chaudoir, 1850* No data Chaudoir, 1850
Ega delicatula (Motschulsky, 1864)* No data Motschulsky, 1864
Euphorticus Horn, 1881 Euphorticus laevicollis (Reiche, 1843) No data Zamorano et al., 2019
Lachnophorus Dejean, 1831 Lachnophorus angusticollis Putzeys, 1878* ANT Putzeys, 1878b
Lachnophorus maculatus Chaudoir, 1850* No data Chaudoir, 1850
Lachnophorus pictipennis Bates, 1871 MAG Martínez, 2003
Lachnophorus sabanillae Liebke, 1936 No data Liebke, 1936
Lachnophorus signatipennis Chaudoir, 1850* No data Chaudoir, 1850
Peruphorticus Erwin & Zamorano, 2014 Peruphorticus pallipes (Reiche, 1843)* No data Zamorano et al., 2019; Reiche, 1843b
Harpalinae Bonelli, 1810 Lachnophorini LeConte, 1853 Peruphorticus Erwin & Zamorano, 2014 Peruphorticus rugatus (Reiche, 1843)* No data Zamorano et al., 2019; Reiche, 1843b
Peruphorticus rugosus (Dejean, 1831)* BOL Zamorano et al., 2019
Stenocheila Laporte de Castelnau, 1832 Stenocheila lacordairei Laporte de Castelnau, 1832 MET Martínez, 2003
Lebiini Bonelli, 1810 Agra Fabricius, 1801 Agra aculeata Chaudoir, 1854* No data Chaudoir, 1854
Agra amoena Chaudoir, 1861* No data Chaudoir, 1861
Agra ardoini Straneo, 1982* CUN Straneo, 1982
Agra aurora Liebke, 1940* No data Liebke, 1940
Agra azurea Chaudoir, 1861* No data Chaudoir, 1861
Agra baleni Steinheil, 1875* BOY Steinheil, 1875a
Agra barrensis Straneo, 1955 No data Erwin, 2002
Agra bogotana Straneo, 1979* CUN Straneo, 1982
Agra castaneipes Bates, 1883 No data Erwin, 2002
Agra cauca Erwin, 1998* CAU Erwin, 1998
Agra cribricollis Chaudoir, 1861* No data Chaudoir, 1861
Agra cyaneucnemes Erwin, 1984* MAG Erwin, 1984
Agra denticulata Steinheil, 1875* ANT Steinheil, 1875a
Agra dorazul Erwin, 1984* SAN Erwin, 1984
Agra eowilsoni Erwin, 1998* VAC Erwin, 1998
Agra incisa Liebke, 1938 MAG Erwin, 2002
Agra jedlickai Liebke, 1938* No data Liebke, 1938
Agra macra Steinheil, 1875* ANT Steinheil, 1875a
Agra magdalena Erwin, 1987* SAN Erwin, 1987
Agra mauritii Straneo, 1982* CUN Straneo, 1982
Agra multifoveolata Steinheil, 1875* ANT Steinheil, 1875a
Agra nova Straneo, 1982* CUN Straneo, 1982
Agra pallipes Liebke, 1938* No data Liebke, 1938
Agra palmata Steinheil, 1875* ANT, BOY, SAN Steinheil, 1875a; Erwin, 1984
Agra pehlkei Liebke, 1938* SAN, TOL Erwin, 1987
Agra perrinae Straneo, 1982* CUN Straneo, 1982
Agra piligera Straneo, 1982* CUN Straneo, 1982
Agra proxima Straneo, 1982* CUN Straneo, 1982
Harpalinae Bonelli, 1810 Lebiini Bonelli, 1810 Agra Fabricius, 1801 Agra setifemoris Straneo, 1982* No data Straneo, 1982
Agra sexdentata Straneo, 1982* VAC Straneo, 1982
Agra smaragdinipennis (Steinheil, 1875)* MAG Erwin, 1983; Steinheil, 1875a
Agra sparsepunctata Straneo, 1982* CUN Straneo, 1982
Agra spinosa Liebke, 1940* No data Liebke, 1940; Mroczkowski, 1960
Agra strangulata Chaudoir, 1863 VAC, CUN Erwin, 1991; Erwin, 2002
Agra subtilicornis Straneo, 1982* CUN Straneo, 1982
Agra tetraspina Straneo, 1982* CUN Straneo, 1982
Agra variabilis Straneo, 1982* CUN Straneo, 1982
Apenes LeConte, 1851 Apenes (Apenes) aerea Steinheil, 1875* CUN Steinheil, 1875a
Apenes (Apenes) cayennensis (Buquet, 1835) No data Motschulsky, 1864
Apenes (Apenes) maculata (Gory, 1833) No data Reiche, 1842b
Apenes (Apenes) mazoreoides Chaudoir, 1875* No data Chaudoir, 1875
Apenes (Apenes) quadripunctata (Reiche, 1842)* No data Reiche, 1842b
Apenes (Apenes) steinheili Ball & Shpeley, 1992* TOL Ball & Shpeley, 1992
Apenes (Apenes) stigmata Liebke, 1939* No data Mroczkowski, 1960
Apenes (Didymochaeta) postica (Dejean, 1831)* BOL Chaudoir, 1875
Axinopalpus LeConte, 1846 Axinopalpus pusillus (Dejean, 1831) BOL Dejean, 1831
Calleida Latreille, 1824 Calleida (Calleida) amethystina (Fabricius, 1787) VAC Arenas-Clavijo & Posso-Gómez, 2017
Calleida (Calleida) bicolor Reiche, 1842 No data Reiche, 1842b
Calleida (Calleida) bogotana Liebke, 1935* CUN Mroczkowski, 1960
Calleida (Calleida) chevrolati Chaudoir, 1873* No data Chaudoir, 1873
Calleida (Calleida) conica Reiche, 1842* No data Reiche, 1842c
Harpalinae Bonelli, 1810 Lebiini Bonelli, 1810 Calleida Latreille, 1824 Calleida (Calleida) cuprea Chaudoir, 1873* No data Chaudoir, 1873
Calleida (Calleida) flava Chevrolat, 1833 No data Reiche, 1842c; Bates, 1883
Calleida (Calleida) fulvipes Reiche, 1842* VAC Reiche, 1842c; Arenas-Clavijo & Posso-Gómez, 2017
Calleida (Calleida) janthina Reiche, 1842* No data Reiche, 1842c
Calleida (Calleida) koppeli Steinheil, 1875* BOY Steinheil, 1875a
Calleida (Calleida) lindigi Chaudoir, 1873* No data Chaudoir, 1873
Calleida (Calleida) lurida Chaudoir, 1873* No data Chaudoir, 1873
Calleida (Calleida) ohausi Liebke, 1939 VAC Arenas-Clavijo & Posso-Gómez, 2017
Calleida (Calleida) onorei Casale, 2008 VAC Arenas-Clavijo & Posso-Gómez, 2017
Calleida (Calleida) pallida Reiche, 1842* No data Reiche, 1842c
Calleida (Calleida) resplendens Reiche, 1842* CUN Reiche, 1842b
Calleida (Calleida) rutilans Chaudoir, 1850* VAC Chaudoir, 1850; Arenas-Clavijo & Posso-Gómez, 2017
Calleida (Calleida) sanguinicollis Dejean, 1831 BOL Dejean, 1831
Calleida (Calleida) schumacheri Steinheil, 1875 ANT, VAC Arenas-Clavijo & Posso-Gómez, 2017; Steinheil 1875a
Calleida (Calleida) similis Reiche, 1842 No data Reiche, 1842c; Bates, 1883
Calleida (Calleida) smaragdinipennis Reiche, 1842* No data Reiche, 1842b
Calleida (Calleida) smaragdula Reiche, 1843* No data Reiche, 1842c
Calleida (Calleida) suturella Reiche, 1842 No data Reiche, 1842b; Chaudoir, 1852
Calleida (Calleida) tibialis Brullé, 1837 No data Chaudoir, 1873
Calleida (Calleida) viridana Liebke, 1939* No data Mroczkowski, 1960
Harpalinae Bonelli, 1810 Lebiini Bonelli, 1810 Calleida Latreille, 1824 Calleida (Callidiola) aurata Motschulsky, 1864 No data Bates, 1883
Callidadelpha Steinheil, 1875 Callidadelpha bogotana Steinheil, 1875* CUN Steinheil, 1875a
Carbonellia Mateu, 1968 Carbonellia (Pseudocarbonellia) atra (Mateu, 1972) NSA Mateu, 1972
Coptodera Dejean, 1825 Coptodera (Coptodera) acutipennis (Buquet, 1835) MAG Shpeley & Ball, 1993
Coptodera (Coptodera) aeneorufa Bates, 1869 AMA Martínez, 2003
Coptodera (Coptodera) apicalis Shpeley & Ball, 1993 VAC Shpeley & Ball, 1993
Coptodera (Coptodera) chalcites Bates, 1869 AMA Shpeley & Ball, 1993; Martínez, 2003
Coptodera (Coptodera) championi Bates, 1883 VAC Shpeley & Ball, 1993
Coptodera (Coptodera) festiva Dejean, 1825 MAG Martínez, 2003
Coptodera (Coptodera) megalops Bates, 1869 CAQ Shpeley & Ball, 1993
Coptodera (Coptodera) nigrostriata (Reiche, 1843) MAG Shpeley & Ball, 1993
Coptodera (Coptodera) nitidula (Buquet, 1835) No data Shpeley & Ball, 1993
Coptodera (Coptodera) picea Dejean, 1826 TOL Shpeley & Ball, 1993
Coptodera (Coptodera) relucens Bates, 1869 AMA Martínez, 2003
Coptodera (Coptodera) schaumii Chaudoir, 1861 CHO, SAN, BOY, CAL, VAC Shpeley & Ball, 1993
Coptodera (Coptodera) transversa (Reiche, 1843) No data Reiche, 1843d; Shpeley & Ball, 1993
Coptodera (Coptodera) undulata Perty, 1830 No data Shpeley & Ball, 1993
Coptodera (Coptodera) versicolor Bates, 1869 AMA Shpeley & Ball, 1993
Cryptobatis Eschscholtz, 1829 Cryptobatis janthoptera (Reiche, 1842) No data Reiche, 1842c; Bates, 1883
Cylindronotum Putzeys, 1845 Cylindronotum nevermanni (Liebke, 1930) MET Erwin, 2004
Dromius Bonelli, 1810 Dromius (Dromius) bordoni Mateu, 1979* CUN Mateu, 1991
Dromius (Dromius) martae Mateu, 1991* MAG Mateu, 1991
Harpalinae Bonelli, 1810 Lebiini Bonelli, 1810 Dromius Bonelli, 1810 Dromius bohumilae Mateu, 1982* No data Mateu, 1991
Dromius columbianus Mateu, 1973* No data Mateu, 1991
Eucheila Dejean, 1829 Eucheila (Inna) boyeri (Solier, 1835) MAG Shpeley & Ball, 2000
Eucheila (Inna) costulata (Chaudoir, 1872) No data Shpeley & Ball, 2000
Euphorticus laevicollis (Reiche, 1843) No data Zamorano et al., 2019
Euplatia Chaudoir, 1872 Euplatia columbica Steinheil, 1875* ANT Steinheil, 1875a
Euproctinus Leng & Mutchler, 1927 Euproctinus (Neoeuproctus) columbianus Shpeley, 1986* No data Shpeley, 1986
Euproctinus (Neoeuproctus) howdeni Shpeley, 1986* VAC Shpeley, 1986
Euproctinus (Neoeuproctus) puzteysi (Chaudoir, 1873)* CUN Shpeley, 1986
Euproctinus (Neoeuproctus) quadriplagiatus (Reiche, 1842) No data Reiche, 1842c; Shpeley, 1986
Eurycoleus Chaudoir, 1848 Eurycoleus fofus Reichardt, 1976 AMA Shpeley & Ball, 2000
Eurycoleus macularius (Chevrolat, 1835) CES, CUN, BOY, VAC Shpeley & Ball, 2000; Ruiz-Tapiador & Arenas, 2015
Hyboptera Chaudoir, 1873 Hyboptera angulicollis Chaudoir, 1873 AMA, CAQ, NAR Erwin & Henry, 2017; Martínez, 2003
Hyboptera auxiliadora Erwin, 2004 BOL Sarmiento-Roa et al., 2020
Hyboptera tiputini Erwin & Henry, 2017 AMA, CHO, VAC Erwin & Henry, 2017
Hyboptera tuberculata (Dejean, 1825) AMA Erwin & Henry, 2017; Martínez, 2003
Hyboptera verrucosa (Reiche, 1842) AMA Erwin & Henry, 2017
Lebia Latreille, 1802 Lebia (Chelonodema) championi (Bates, 1883) CUN Reichardt, 1972
Lebia (Chelonodema) erotyloides Reichardt, 1972 COR Martínez, 2003
Lebia (Chelonodema) howdeni Reichardt, 1972* VAC Reichardt, 1972
Lebia (Chelonodema) ocelligera (Bates, 1883) CUN, BOY Reichardt, 1972
Harpalinae Bonelli, 1810 Lebiini Bonelli, 1810 Lebia Latreille, 1802 Lebia (Lebia) amabilis (Chaudoir, 1871) No data Bates, 1883
Lebia (Lebia) amoenula (Chaudoir, 1871) No data Chaudoir, 1871
Lebia (Lebia) angulata Dejean, 1831 BOL Dejean, 1831
Lebia (Lebia) atricapillus Liebke, 1931* SAN Putzeys, 1878b
Lebia (Lebia) bitaeniata Chevrolat, 1834 AMA, VAC Martínez, 2003
Lebia (Lebia) brachinoides Reiche, 1842 No data Reiche, 1842c; Bates, 1883
Lebia (Lebia) callida Liebke, 1938* No data Mroczkowski, 1960
Lebia (Lebia) cannae Steinheil, 1875* CAL, CAQ, VID Steinheil 1875a
Lebia (Lebia) chlorotica Dejean, 1831 No data Bates 1883,
Lebia (Lebia) cognata Chaudoir, 1871* No data Chaudoir, 1871
Lebia (Lebia) confusula Chaudoir, 1871* BOL Chaudoir, 1871
Lebia (Lebia) contigua Chaudoir, 1871* No data Chaudoir, 1871
Lebia (Lebia) dentata Chaudoir, 1871* PUT Chaudoir, 1870
Lebia (Lebia) discernenda Chaudoir, 1871* No data Chaudoir, 1870
Lebia (Lebia) goudoti (Chaudoir, 1871)* No data Chaudoir, 1871
Lebia (Lebia) gratiosa (Chaudoir, 1871)* No data Chaudoir, 1871
Lebia (Lebia) haplomera Chaudoir, 1871* No data Chaudoir, 1871
Lebia (Lebia) lacerata Chaudoir, 1871* No data Chaudoir, 1870
Lebia (Lebia) leptodera (Chaudoir, 1871)* No data Chaudoir, 1871
Lebia (Lebia) longiloba Chaudoir, 1871* BOL Bates, 1883; Chaudoir, 1871
Lebia (Lebia) melanoptera Chaudoir, 1871* No data Chaudoir, 1871
Lebia (Lebia) mirabilis Bates, 1883 CAQ Martínez, 2003
Lebia (Lebia) myops Dejean, 1831* BOL Chaudoir, 1871
Harpalinae Bonelli, 1810 Lebiini Bonelli, 1810 Lebia Latreille, 1802 Lebia (Lebia) nigrolineata Reiche, 1842* BOL Reiche, 1842c
Lebia (Lebia) nigromaculata Gory, 1833 BOL Chaudoir, 1870
Lebia (Lebia) pallipes Gory, 1833 No data Reiche, 1842c
Lebia (Lebia) picta (Steinheil, 1875)* BOY Steinheil, 1875a
Lebia (Lebia) puella Dejean, 1831* BOL Chaudoir, 1870
Lebia (Lebia) reflexicollis Chaudoir, 1843 No data Chaudoir, 1871
Lebia (Lebia) ruficeps (Chaudoir, 1871)* No data Chaudoir, 1871
Lebia (Lebia) rugifrons Dejean, 1831* BOL Chaudoir, 1870
Lebia (Lebia) smaragdinipennis Reiche, 1842* No data Reiche, 1842c
Lebia (Lebia) subtilis (Chaudoir, 1871) No data Chaudoir, 1871
Lebia (Lebia) terminalis Putzeys, 1846 No data Chaudoir, 1871
Lebia (Lebia) variegata Dejean, 1831* BOL Dejean, 1831
Lebia (Lebia) vicina (Chaudoir, 1871) MAG Chaudoir, 1871; Bates, 1883
Lebia argutula (Chaudoir, 1871)* No data Chaudoir, 1871
Lebia disconotata (Chaudoir, 1871)* BOL Chaudoir, 1871
Lebia distinguenda Putzeys, 1846 No data Chaudoir, 1870
Lebia heydenii Putzeys, 1845* No data Chaudoir, 1870
Lebia limbata Steinheil, 1875* QUI Steinheil, 1875a
Lelis Chaudoir, 1869 Lelis quadrisignata (Buquet, 1835) AMA Shpeley & Ball, 2000
Lelis rutila (Bates, 1869) BOY Shpeley & Ball, 2000; Steinheil, 1875a
Mochtherus Schmidt-Göbel, 1846 Mochtherus tetraspilotus (MacLeay, 1825) VAC Torres-Dominguez et al., 2020
Negrea Mateu, 1968 Negrea opaca Mateu, 1982 MAG Martínez, 2003
Negrea scutellaris (Dejean, 1834) BOL Mateu, 1982a
Harpalinae Bonelli, 1810 Lebiini Bonelli, 1810 Nemotarsus LeConte, 1853 Nemotarsus fallax (Dejean, 1831) BOL Bates, 1883
Onota Chaudoir, 1873 Onota angulicollis (Reiche, 1842) No data Reiche, 1842c; Erwin, 2004
Onota rutilans Chaudoir, 1873 AMA Martínez, 2003
Phloeoxena Chaudoir, 1870 Phloeoxena (Phloeoxena) biundata Steinheil, 1875* NSA Shpeley & Ball, 2000; Steinheil, 1875b
Phloeoxena (Oenaphelox) signata Dejean, 1825 No data Chaudoir, 1870
Plochionus Dejean, 1821 Plochionus (Menidius) faviger Chaudoir, 1873* No data Chaudoir, 1873
Plochionus (Menidius) pictipennis (Reiche, 1842)* VID Reiche, 1842c
Plochionus (Menidius) pictus Chaudoir, 1873* No data Chaudoir, 1873
Plochionus (Plochionus) pallens (Fabricius, 1775) No data Bates, 1883; Chaudoir, 1872
Pseudotoglossa Mateu, 1961 Pseudotoglossa inaequalis (Chaudoir, 1873) AMA Martínez, 2003
Pseudotoglossa obscurella (Bates, 1878) MAG Erwin, 2004; Mateu, 1961
Pseudotoglossa terminalis (Chaudoir, 1873) No data Erwin, 2004; Mateu 1961
Stenognathus Chaudoir, 1843 Stenognathus (Gnathostenus) dentifemoratus Shpeley & Ball, 2000* No data Shpeley & Ball, 2000
Stenognathus (Pristolomus) dentifer (Chaudoir, 1870) No data Shpeley & Ball, 2000
Stenognathus (Stenognathus) crenulatus Chaudoir, 1870 CUN, PUT, VAC Shpeley & Ball, 2000
Stenognathus (Stenognathus) longipennis Chaudoir, 1877* BOY, VAC Shpeley & Ball, 2000
Stenognathus (Stenognathus) platypterus Chaudoir, 1870 AMA Shpeley & Ball, 2000
Stenognathus (Stenognathus) procerus (Putzeys, 1878)* ANT Putzeys, 1878b; Shpeley & Ball, 2000
Thoasia Liebke, 1939 Thoasia rugifrons Liebke, 1939 MAG Reichardt, 1968
Morionini Brullé, 1834 Morion Latreille, 1810 Morion cordatus Chaudoir, 1837 AMA, CES, CHO, NAR Reiche, 1843a; Martínez, 2003
Morion cyclomus Chaudoir, 1854 No data Chaudoir, 1854
Harpalinae Bonelli, 1810 Morionini Brullé, 1834 Morion Latreille, 1810 Morion monilicornis (Latreille, 1805) No data Chaudoir, 1854; Bates, 1883
Morion simplex Dejean, 1826 MAG Martínez, 2003
Moriosomus Motschulsky, 1855 Moriosomus seticollis Straneo, 1985 No data Erwin & Moore, 2007
Odacanthini Laporte de Castelnau, 1834 Calophaena Klug, 1821 Calophaena acuminata (Olivier, 1790) No data Reiche, 1842a
Calophaena arcuata (Guérin-Méneville, 1844) No data Bates, 1883
Calophaena bicincta (Dejean & Boisduval, 1829) No data Motschulsky, 1864
Calophaena cincta (Gray, 1832) No data Steinheil, 1875a
Calophaena grandispina Liebke, 1930 No data Mroczkowski, 1960
Calophaena laevigata Bates, 1878 CHO Martínez, 2003
Calophaena lafertei Guérin-Méneville, 1844* No data Guérin-Méneville, 1844
Calophaena unifasciata Chaudoir, 1861 No data Chaudoir, 1861
Calophaena virgata Liebke, 1938* No data Mroczkowski, 1960
Colliuris DeGeer, 1774 Colliuris (Apiodera) funckii (Putzeys, 1845) VID Martínez, 2003
Colliuris (Apiodera) rugicollis (Dejean, 1825) No data Chaudoir, 1863
Colliuris (Apiodera) subdistincta (Chaudoir, 1863) AMA Martínez, 2003
Colliuris (Apioderella) rudis (Chaudoir, 1872) RIS Martínez, 2003
Colliuris (Colliurella) amoena (Chaudoir, 1863)* No data Chaudoir, 1863
Colliuris (Colliurita) variolosa (Chaudoir, 1863)* No data Chaudoir, 1863
Colliuris (Mimocasnonia) fusca (Reiche, 1842) No data Reiche, 1842a
Colliuris (Plagiorhytis) corrusca (Chaudoir, 1863) No data Chaudoir, 1863
Colliuris (Plagiorhytis) plicaticollis (Reiche, 1842) No data Reiche, 1842a
Colliuris (Pseudocasnonia) viridicollis (Chaudoir, 1863) No data Chaudoir, 1863
Colliuris tetrastigma (Chaudoir, 1863) AMA Martínez, 2003
Oodinus Motschulsky, 1864 Oodinus alutaceus (Bates, 1882) AMA Martínez, 2003
Harpalinae Bonelli, 1810 Odacanthini Laporte de Castelnau, 1834 Pentagonica Shmidt-Göbel, 1846 Pentagonica flavipes (LeConte, 1853) AMA, MAG, NAR Reichardt, 1968; Martínez, 2003
Pentagonica maculicornis Bates, 1883 MAG Reichardt, 1968
Pentagonica ochracea Reichardt, 1968 VID Martínez, 2003
Pentagonica roedingeri Liebke, 1951 AMA Martínez, 2003
Pentagonica scutellaris Chaudoir, 1877 AMA Martínez, 2003
Pentagonica trivittata (Dejean, 1831) BOL Dejean, 1831
Stenocrepis Chaudoir, 1857 Stenocrepis (Stenocrepis) pauper Chaudoir, 1857 No data Chaudoir, 1857
Stenocrepis (Stenous) fuscipes (Laferté-Sénectère, 1851) No data Laferté-Sénectère, 1851
Stenocrepis (Stenous) gilvipes (Laferté-Sénectère, 1851) No data Reiche, 1843b; Chaudoir, 1857
Stenocrepis (Stenous) metallica (Dejean, 1826) No data Chaudoir; 1857
Stenocrepis (Stenous) tibialis (Chevrolat, 1834) No data Laferté-Sénectère, 1851
Stenocrepis aeruginea (Laferté-Sénectère, 1851) No data Laferté-Sénectère, 1851
Stenocrepis flavicrus (Laferté-Sénectère, 1851)* No data Laferté-Sénectère, 1851
Peleciini Chaudoir, 1880 Pelecium Kirby, 1819 Pelecium (Pelecidium) sulcatum Guérin-Méneville, 1843* MAG, NSA, VAC Straneo & Ball, 1989
Pelecium laevigatum Guérin-Méneville, 1843* TOL Straneo & Ball, 1989
Perigonini Horn, 1881 Mizotrechus Bates, 1872 Mizotrechus gorgona Erwin, 2011* CAU Erwin, 2011
Perigona Laporte de Castelnau, 1835 Perigona (Perigona) columbiana Putzeys, 1878* BOY Putzeys, 1878b
Platynini Bonelli, 1810 Dyscolus Dejean, 1831 Dyscolus (Dyscolidion) bracteatus (Moret, 1990) CAU Arenas-Clavijo & González, 2018
Dyscolus (Dyscolidion) cyanonotus Chaudoir, 1850 No data Chaudoir, 1850; Martínez, 2003
Dyscolus (Dyscolidion) laticollis (Reiche, 1843)* No data Martínez & Ball, 2003
Dyscolus (Dyscolidion) leptomorphus (Chaudoir, 1879)* CUN Martínez & Ball, 2003
Dyscolus (Dyscolidion) politus (Putzeys, 1878) CUN, ANT Putzeys, 1878
Dyscolus (Dyscolidion) princeps (Bates, 1878) No data Martínez & Ball, 2003
Harpalinae Bonelli, 1810 Platynini Bonelli, 1810 Dyscolus Dejean, 1831 Dyscolus (Dyscolidion) punctatostriatus (Putzeys, 1878)* BOY, TOL Martínez & Ball, 2003
Dyscolus (Dyscolidion) sinuosus (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolidion) viridiauratus (Bates, 1878) No data Martínez & Ball, 2003
Dyscolus (Dyscolus) acutipennis Chaudoir, 1850* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) acutus (Putzeys, 1878)* NSA Martínez & Ball, 2003
Dyscolus (Dyscolus) agonoides (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) ahenonotus (Putzeys, 1878)* CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) alpaeoides (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) anthracinus (Putzeys, 1878)* CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) asphaltinus (Chaudoir, 1878) CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) atratus (Chaudoir, 1859) MAG Martínez & Ball, 2003
Dyscolus (Dyscolus) atroaeneus (Putzeys, 1878)* BOY Martínez & Ball, 2003; Perrault, 1990
Dyscolus (Dyscolus) beryllinus (Putzeys, 1878)* CAL Martínez & Ball, 2003
Dyscolus (Dyscolus) bispinosus (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) bogotensis Perrault, 1992* CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) brachypterus (Chaudoir, 1859)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) brevipennis (Motschulsky, 1865)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) brevis (Putzeys, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) carbonarius (Putzeys, 1878)* CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) caucaensis Perrault, 1992* VAC Martínez & Ball, 2003
Dyscolus (Dyscolus) chalconotus (Chaudoir, 1878)* CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) ciliatus (Chaudoir, 1878)* CUN Martínez & Ball, 2003
Harpalinae Bonelli, 1810 Platynini Bonelli, 1810 Dyscolus Dejean, 1831 Dyscolus (Dyscolus) clarus (Chaudoir, 1878)* BOY Martínez & Ball, 2003
Dyscolus (Dyscolus) conicus (Chaudoir, 1879) MAG Martínez & Ball, 2003
Dyscolus (Dyscolus) consanguineus (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) cordicollis (Motschulsky, 1865) No data Martínez & Ball, 2003
Dyscolus (Dyscolus) corvinus (Dejean, 1831)* CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) cyaneocupreus (Putzeys, 1878)* ANT, CAL, TOL Chaudoir, 1878; Martínez & Ball, 2003
Dyscolus (Dyscolus) cyanicollis Brullé, 1834* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) despiciendus (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) donrwi Perrault, 1993* BOY, CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) dyschromus (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) feronioides (Reiche, 1843)* ARA, BOY, CUN, MAG, MET, QUI, SAN, VAU Martínez & Ball, 2003; Camero, 2003
Dyscolus (Dyscolus) grandicollis (Reiche, 1843)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) hexacoelus (Chaudoir, 1879) No data Chaudoir, 1879
Dyscolus (Dyscolus) inconspicuus (Chaudoir, 1878)* CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) interruptus (Putzeys, 1878)* ANT Martínez & Ball, 2003
Dyscolus (Dyscolus) isabellae Camero, 2010* MAG Camero, 2010
Dyscolus (Dyscolus) kennedyensis Camero, 2010* MAG Camero, 2010
Dyscolus (Dyscolus) landolti (Putzeys, 1878)* NSA, SAN Martínez & Ball, 2003
Dyscolus (Dyscolus) latidens (Chaudoir, 1859) No data Martínez & Ball, 2003
Dyscolus (Dyscolus) longipennis (Reiche, 1843)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) lucidus (Chaudoir, 1879)* BOL Martínez & Ball, 2003
Dyscolus (Dyscolus) melas (Putzeys, 1878)* SAN Martínez & Ball, 2003
Harpalinae Bonelli, 1810 Platynini Bonelli, 1810 Dyscolus Dejean, 1831 Dyscolus (Dyscolus) micans (Putzeys, 1878)* BOL, SAN Martínez & Ball, 2003
Dyscolus (Dyscolus) monterredonda Perrault, 1992* CUN, HUI Martínez & Ball, 2003
Dyscolus (Dyscolus) morosus (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) muzo Perrault, 1993* CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) obesulus (Chaudoir, 1878)* MAG Martínez & Ball, 2003
Dyscolus (Dyscolus) oopterus (Chaudoir, 1859)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) ovatus (Putzeys, 1878)* BOY Martínez & Ball, 2003
Dyscolus (Dyscolus) phaeocnemis (Chaudoir, 1879)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) physopterus (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) piceolus (Chaudoir, 1878)* SAN Martínez & Ball, 2003
Dyscolus (Dyscolus) platynoides (Chaudoir, 1878)* ARA, BOY, CAL, CUN, MAG, MET, QUI, SAN, TOL Martínez & Ball, 2003; Camero, 2003
Dyscolus (Dyscolus) protensus (Putzeys, 1878)* CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) pseudoconicus Perrault, 1992* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) purpuratus Reiche, 1843 ANT, CAL, RIS, VAC Martínez & Ball, 2003
Dyscolus (Dyscolus) quadricollis (Chaudoir, 1859)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) reichei (Perrault, 1989) No data Martínez & Ball, 2003
Dyscolus (Dyscolus) rutilans (Motschulsky, 1865) BOY Martínez & Ball, 2003
Dyscolus (Dyscolus) seriepunctatus (Chaudoir, 1859)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) spinipennis (Reiche, 1843)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) steinheili Perrault, 1993* ARA, BOY, CAL, CUN, MAG, MET, NSA, SAN, VAU Martínez & Ball, 2003; Camero, 2003
Dyscolus (Dyscolus) striatulus (Chaudoir, 1878)* No data Martínez & Ball, 2003
Harpalinae Bonelli, 1810 Platynini Bonelli, 1810 Dyscolus Dejean, 1831 Dyscolus (Dyscolus) subangulatus (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) subiridescens (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) subreflexus (Chaudoir, 1878)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) subviolaceus (Chaudoir, 1842) RIS, SAN Martínez & Ball, 2003
Dyscolus (Dyscolus) sulcatus (Guérin-Méneville, 1844)* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) thiemei Perrault, 1990* No data Martínez & Ball, 2003
Dyscolus (Dyscolus) trapezicollis Chaudoir, 1878 CUN, NSA, SAN Martínez & Ball, 2003; Perrault, 1990
Dyscolus (Dyscolus) unipunctatus Perrault, 1990* CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) vegaensis Perrault, 1992* CUN Martínez & Ball, 2003
Dyscolus (Dyscolus) villavicencio Perrault, 1992* MET Martínez & Ball, 2003
Dyscolus (Glyptolenopsis) aeneipennis (Dejean, 1831) BOL Dejean, 1831
Dyscolus (Glyptolenopsis) degallieri (Perrault, 1991) NAR Martínez & Ball, 2003
Dyscolus (Stenocnemion) arenasi Moret, 2019* VAC Moret, 2019
Dyscolus (Stenocnemion) martinezae Moret, 2019* VAC Moret, 2019
Glyptolenoides Perrault, 1991 Glyptolenoides azureipennis (Chaudoir, 1859) No data Martínez & Ball, 2003
Glyptolenoides azureus (Chaudoir, 1859) NSA Martínez & Ball, 2003
Glyptolenoides cyclothorax (Chaudoir, 1879) No data Martínez & Ball, 2003
Glyptolenoides elegantulus (Chaudoir, 1878)* No data Martínez & Ball, 2003
Glyptolenoides purpuripennis (Chaudoir, 1879) No data Perrault, 1991
Glyptolenoides sulcipennis (Chaudoir, 1879)* CUN Martínez & Ball, 2003
Glyptolenoides sulcitarsis (Chaudoir, 1878)* No data Martínez & Ball, 2003
Glyptolenus Bates, 1878 Glyptolenus apicestriatus (Reiche, 1843)* No data Martínez & Ball, 2003
Harpalinae Bonelli, 1810 Platynini Bonelli, 1810 Glyptolenus Bates, 1878 Glyptolenus chalybeus (Dejean, 1831) ARA, BOY, CAL, CAU, CUN, MET, SAN, TOL, VAU Camero, 2003; Arenas-Clavijo & González, 2018
Glyptolenus convexiusculus (Chaudoir, 1878) No data Chaudoir, 1878
Glyptolenus janthinus (Dejean, 1831) No data Martínez & Ball, 2003
Glyptolenus nigrita (Chaudoir, 1879)* No data Martínez & Ball, 2003
Glyptolenus nitidipennis (Chaudoir, 1850)* No data Martínez & Ball, 2003
Glyptolenus spinosus (Reiche, 1843)* No data Martínez & Ball, 2003
Incagonum Liebherr, 1994 Incagonum aeneum (Reiche, 1843) BOY, CAU, HUI, CAL, CUN, QUI Martínez & Ball, 2003
Incagonum pedestre (Putzeys, 1878)* CUN Martínez & Ball, 2003
Sericoda Kirby, 1837 Sericoda bembidioides Kirby, 1837 BOY, CUN, TOL Martínez & Ball, 2003
Pterostichini Bonelli, 1810 Abaris Dejean, 1831 Abaris (Abaridius) retiaria Will, 2002 MAG Will, 2002
Abaris (Abaris) aenea Dejean, 1831 BOL, MAG Will, 2002
Abaris (Abaris) basistriata Chaudoir, 1874 MAG Will, 2002
Blennidus Motschulsky, 1865 Blennidus aberrans (Straneo, 1985)* MAG Straneo, 1985
Blennidus angularis (Straneo, 1985)* MAG Straneo, 1985
Blennidus bistriatus (Straneo, 1951)* MAG Straneo, 1951
Blennidus darlingtoni (Straneo, 1951)* MAG Straneo, 1951
Blennidus davidsoni (Straneo, 1985)* MAG Straneo, 1985
Blennidus dianae Camero, 2006* MAG Camero, 2006
Blennidus kochalkai (Straneo, 1985)* MAG Straneo, 1985
Blennidus laevigatus (Straneo, 1951)* MAG Straneo, 1951
Blennidus laevis (Straneo, 1951) MAG Straneo, 1951
Blennidus minutus (Straneo, 1951)* MAG Straneo, 1951
Harpalinae Bonelli, 1810 Pterostichini Bonelli, 1810 Blennidus Motschulsky, 1865 Blennidus montanus (Straneo, 1951)* MAG Straneo, 1951
Blennidus parvulus (Straneo, 1951)* MAG Straneo, 1951
Blennidus smaragdinus (Straneo, 1951)* MAG Straneo, 1951
Blennidus striolatus (Straneo, 1951)* MAG Straneo, 1951
Blennidus subcordatus (Straneo, 1951)* MAG Straneo, 1951
Blennidus uniformis (Straneo, 1951)* MAG Straneo, 1951
Hybothecus Chaudoir, 1874 Hybothecus incrassatus Chaudoir, 1874* No data Chaudoir, 1874b
Hybothecus sculptilis (Putzeys, 1878)* ANT Putzeys, 1878b
Loxandrus LeConte, 1852 Loxandrus latifascia Straneo, 1991 No data Straneo, 1991
Loxandrus minimus Straneo, 1951 MAG Straneo, 1951
Loxandrus interruptus Tschitscherine, 1900* TOL Tschitschérine, 1900
Loxandrus opaculus Bates, 1871 CUN Martínez, 2003
Loxandrus ornatus Putzeys, 1878* CUN Putzeys, 1878b; Straneo, 1991
Loxandrus pictoides Straneo, 1991 VAC Straneo, 1991; Martínez, 2003
Loxandrus tetrastigma Bates, 1871 RIS, VAC Arenas-Clavijo & Posso-Gómez, 2017
Oribazus Chaudoir, 1874 Oribazus catenulatus Chaudoir, 1874 No data Chaudoir, 1874b
Oribazus quinquestriatus Chaudoir, 1874 No data Chaudoir, 1874b
Oxycrepis Oxycrepis leucocera Reiche, 1843 No data Will, 2005
Pseudabarys Pseudabarys lebasi (Chaudoir, 1874)* No data Chaudoir, 1874a
Stolonis Motschulsky, 1866 Stolonis elegans (Dejean, 1831)* BOL Chaudoir, 1873
Stolonis parvula (Straneo, 1951)* MAG Will, 2005
Sphodrini Laporte de Castelnau, 1834 Laemostenus Bonelli, 1810 Laemostenus (Laemostenus) complanatus (Dejean, 1828) AMA, BOY, CAL, CUN, HUI, MET, NAR, NSA, RIS, SAN, TOL Martínez & Ball, 2003
Harpalinae Bonelli, 1810 Zuphiini Bonelli, 1810 Pseudaptinus Laporte de Castelnau, 1834 Pseudaptinus (Thalpius) intermedius (Chaudoir, 1872)* No data Chaudoir, 1872
Zuphioides Ball & Shpeley, 2013 Zuphioides capitum (Liebke, 1933)* No data Liebke, 1933; Ball & Shpeley, 2013
Zuphioides columbianum (Chaudoir, 1872)* No data Chaudoir, 1872; Ball & Shpeley, 2013
Zuphioides exiguum (Putzeys, 1878)* VAC Putzeys, 1878b; Ball & Shpeley, 2013
Licininae Bonelli, 1810 Chlaeniini Brullé, 1834 Chlaenius Bonelli, 1810 Chlaenius (Chlaenius) fallax (Olivier, 1795) No data Chaudoir, 1873
Chlaenius (Chlaenius) leucoscelis Chevrolat, 1835 No data Chaudoir, 1873
Chlaenius (Chlaenius) viridicollis Reiche, 1843 No data Reiche, 1843b
Melaeninae Alluaud, 1934 Melaenini Csiki, 1933 Cymbionotum Baudi di Selve, 1864 Cymbionotum (Procoscinia) fernandezi Ball & Shpeley, 2005* BOL Ball & Shpeley, 2005
Paussinae Latreille, 1807 Ozaenini Hope, 1838 Ozaena Olivier, 1811 Ozaena dentipes Olivier, 1811 MET Martínez, 2003
Ozaena martinezi Ogueta, 1965* MET Martínez, 2003
Pachyteles Perty, 1830 Pachyteles angustatus Chaudoir, 1868* No data Chaudoir, 1868a
Pachyteles baleni Steinheil, 1875* ANT Steinheil, 1875b
Pachyteles castaneus (Dejean, 1831) BOL Chaudoir, 1868a
Pachyteles gyllenhalii (Dejean, 1825) No data Chaudoir, 1852
Pachyteles haroldi Steinheil, 1875* BOY Steinheil, 1875b
Pachyteles politus (Reiche, 1843)* No data Reiche, 1843a; Chaudoir, 1854
Pachyteles seriepunctatus Chaudoir, 1868* No data Chaudoir, 1868
Pachyteles striola Perty, 1830 No data Chaudoir, 1852
Physea Brullé, 1835 Physea testudinea (Klug, 1834) BOY, MET Martínez, 2003
Paussini Latreille, 1806 Homopterus Westwood, 1841 Homopterus cunctans Reichensperger, 1938 CAU Arenas-Clavijo & González, 2018
Homopterus steinbachi Kolbe, 1920 BOY Darlington, 1950
Paussinae Latreille, 1807 Paussini Latreille, 1806 Homopterus Westwood, 1841 Homopterus subcordatus Darlington, 1950 CAQ Sarmiento-Roa et al., 2020
Rhysodinae Laporte de Castelnau, 1840 Rhysodini Laporte de Castelnau, 1840 Clinidium Kirby, 1830 Clinidium (Clinidium) cavicolle Chevrolat, 1873 NSA Bell & Bell, 1985
Clinidium (Clinidium) curvatum Bell & Bell, 1985* NSA Bell & Bell. 1985
Clinidium (Clinidium) granatense Chevrolat, 1873* ANT, CUN, BOY Bell & Bell, 1985; Bell & Bell, 2009
Clinidium (Clinidium) hammondi Bell & Bell, 1985* CUN Bell & Bell, 1985
Clinidium (Clinidium) humboldti Bell & Bell, 1985* No data Bell & Bell, 1985
Clinidium (Clinidium) humile Bell & Bell, 1985* No data Bell & Bell, 1985
Clinidium (Clinidium) integrum Grouvelle, 1903 AMA Bell & Bell, 2009
Clinidium (Clinidium) kochalkai Bell & Bell, 1985* MAG Bell & Bell, 1985
Clinidium (Clinidium) mathani Grouvelle, 1903 CAQ, CAU Bell & Bell, 2009; Martínez, 2003
Clinidium (Clinidium) oberthueri Grouvelle, 1903 NAR Bell & Bell, 2009
Clinidium (Clinidium) penicillatum Bell & Bell, 1985* VAC Bell & Bell, 1985
Clinidium (Clinidium) spatulatum Bell & Bell, 1985 CAU Bell & Bell, 2009; Martínez, 2003
Scaritinae Bonelli, 1810 Clivinini Rafinesque, 1815 Ardistomis Putzeys, 1846 Ardistomis dubia Putzeys, 1846 No data Putzeys, 1846
Ardistomis dyschirioides Putzeys, 1846 No data Putzeys, 1846
Ardistomis ovata Putzeys, 1846* No data Putzeys, 1846
Ardistomis seriepunctata (Brullé, 1843) No data Valdés, 2009
Aspidoglossa Putzeys, 1846 Aspidoglossa mexicana (Chaudoir, 1837) No data Putzeys, 1867
Aspidoglossa pallida Putzeys, 1846* No data Putzeys, 1846
Aspidoglossa rivalis Putzeys, 1846 No data Putzeys, 1846
Aspidoglossa submetallica Putzeys, 1846 No data Putzeys, 1846
Clivina Latreille,1802 Clivina (Clivina) erythropus Putzeys, 1846* No data Putzeys, 1846
Clivina (Clivina) lebasii Putzeys, 1846* No data Putzeys, 1846
Clivina (Clivina) quadrata Putzeys, 1867* BOL Putzeys, 1867
Scaritinae Bonelli, 1810 Clivinini Rafinesque, 1815 Clivina Latreille,1802 Clivina (Paraclivina) fasciata Putzeys, 1846 No data Putzeys, 1846
Clivina (Paraclivina) fuscipes Putzeys, 1846* No data Putzeys, 1846
Clivina (Paraclivina) tristis Putzeys, 1846* No data Putzeys, 1846
Clivina (Paraclivina) tuberculata Putzeys, 1846* CUN Putzeys, 1846
Clivina (Semiclivina) armata Putzeys, 1846* BOL Putzeys, 1846
Clivina (Semiclivina) columbica Putzeys, 1846* No data Putzeys, 1846
Clivina (Semiclivina) dissimilis Putzeys, 1846* No data Putzeys, 1846
Clivina (Semiclivina) laticeps Putzeys, 1846* No data Putzeys, 1846
Clivina (Semiclivina) latimanus Putzeys, 1846* No data Putzeys, 1846
Clivina (Semiclivina) oblita Putzeys, 1867* No data Putzeys, 1867
Cryptomma Putzeys, 1846 Cryptomma multistriatum Putzeys, 1846* No data Putzeys, 1846
Oxydrepanus Putzeys, 1867 Oxydrepanus ovalis Putzeys, 1867* No data Putzeys, 1867
Pyramis Putzeys, 1846 Pyramis crassicornis Putzeys, 1846* No data Putzeys, 1846
Schizogenius Putzeys, 1846 Schizogenius (Genioschizus) impressicollis Putzeys, 1846* No data Whitehead, 1972; Putzeys, 1846
Schizogenius (Genioschizus) impuncticollis Whitehead, 1972* MAG Whitehead, 1972
Schizogenius (Genioschizus) suturalis Whitehead, 1972* MAG Whitehead, 1972
Schizogenius (Genioschizus) szekessyi Kult, 1950 MAG Whitehead, 1972
Schizogenius (Genioschizus) tenuis Bates, 1881 MAG, TOL Whitehead, 1972
Schizogenius (Listropus) dyschirioides Putzeys, 1861 MAG Whitehead, 1972
Schizogenius (Schizogenius) capitalis Putzeys, 1861* ANT Whitehead, 1972
Schizogenius (Schizogenius) darlingtoni Kult, 1950 No data Whitehead, 1972
Schizogenius (Schizogenius) gracilis Putzeys, 1846 No data Putzeys, 1846
Scaritinae Bonelli, 1810 Clivinini Rafinesque, 1815 Schizogenius Putzeys, 1846 Schizogenius (Schizogenius) interstriatus Putzeys, 1878* ANT Putzeys, 1878b
Schizogenius (Schizogenius) pygmaeus Van Dyke, 1925 MAG Whitehead, 1972
Schizogenius (Schizogenius) riparius Putzeys, 1878* TOL Putzeys, 1878b
Schizogenius (Schizogenius) strigicollis Putzeys, 1846* MAG Whitehead, 1972; Putzeys, 1846
Schizogenius (Schizogenius) sulcatulus Putzeys, 1846* No data Putzeys, 1846
Semiardistomis Kult, 1950 Semiardistomis cordicollis (Putzeys, 1846)* No data Valdés, 2012
Semiardistomis pallipes (Dejean, 1831) BOL Dejean, 1831; Valdés, 2012
Forcipatorini Bänninger, 1937 Camptodontus Dejean, 1826 Camptodontus falcatus Putzeys, 1861* No data Putzeys, 1867
Camptodontus reichei Putzeys, 1861* No data Putzeys, 1867
Forcipator Maindron, 1904 Forcipator putzeysii (Chaudoir, 1868)* No data Chaudoir, 1868b
Stratiotes Putzeys, 1846 Stratiotes batesi Putzeys, 1867 NAR Martínez, 2003
Scaritini Bonelli, 1810 Distichus Motschulsky, 1858 Distichus (Lophogenius) orientalis (Bonelli, 1813) No data Bänninger, 1938
Distichus (Lophogenius) lacordairei (Dejean, 1831) No data Bänninger, 1938
Glyptogrus Bates, 1881 Glyptogrus molopinus (Perty, 1830) No data Chaudoir, 1879
Scarites Fabricius, 1775 Scarites (Scarites) heterogrammus Perty, 1830 No data Bänninger, 1938
Scarites (Scarites) illustris Chaudoir, 1880 No data Bänninger, 1938
Scarites (Scarites) zambo Steinheil, 1875* No data Steinheil, 1875b; Bänninger, 1938
Taeniolobus Chaudoir, 1855 Taeniolobus (Taeniolobus) gratus (Chaudoir, 1855)* No data Chaudoir, 1880; Bänninger, 1938
Taeniolobus (Taeniolobus) guerini (Chaudoir, 1855) No data Chaudoir, 1855; Bänninger, 1938
Taeniolobus (Taeniolobus) oberthueri Bänninger, 1938* ANT Steinheil, 1875b; Bänninger, 1938
Taeniolobus (Taeniolobus) thiemei (Bänninger, 1933)* No data Bänninger, 1938
Taeniolobus (Taeniolobus) lebasii (Chaudoir, 1855)* No data Chaudoir, 1855; Bänninger, 1938
Siagoninae Bonelli, 1813 Siagonini Bonelli, 1813 Enceladus Bonelli, 1813 Enceladus gygas Bonelli, 1813 LAG Arenas-Clavijo & Posso-Gómez, 2017
Trechinae Bonelli, 1810 Bembidiini Stephens, 1827 Bembidion Latreille, 1802 Bembidion (Antiperyphanes) angulicolle (Putzeys, 1878)* CUN Putzeys, 1878b
Bembidion (Ecuadion) boyaca Toledano, 2008* BOY Toledano, 2008
Bembidion (Ecuadion) chimborazonum Bates, 1871 VAC Sarmiento-Roa et al., 2020
Bembidion (Ecuadion) cocuyanum Toledano, 2008* BOY Toledano, 2008
Bembidion (Ecuadion) colombianum Toledano, 2008* CUN Toledano, 2008
Bembidion (Ecuadion) putzeysii Csiki, 1928* CUN Putzeys, 1878b
Bembidion (Ecuadion) quebrada Toledano, 2008* TOL Toledano, 2008
Bembidion (Ecuadion) sanctaemarthae Darlington, 1934 MAG Toledano, 2008
Bembidion (Ecuadion) subapterum Darlington, 1934 MAG Toledano, 2008
Bembidion (Notaphus) basiplagiatum (Putzeys, 1878) CUN Putzeys, 1878b
Bembidion (Notaphus) cayambense Bates, 1891 CUN Toledano, 2008
Bembidion (Notaphus) commissum Erichson, 1847 VAC Toledano, 2008
Elaphropus Motschulsky, 1839 Elaphropus (Ammotachys) marchantarius Boyd & Erwin, 2016 AMA Boyd & Erwin, 2016
Elaphropus (Nototachys) occidentalis Boyd & Erwin, 2016 AMA Boyd & Erwin, 2016
Elaphropus (Tachyura) yunax (Darlington, 1939) MAG Martínez, 2003
Erwiniana Paulsen & Smith, 2003 Erwiniana alticola (Erwin, 1994)* CUN Erwin, 1994
Erwiniana anchicaya (Erwin, 1994)* CAU, VAC Erwin, 1994
Erwiniana depressisculptilis (Erwin, 1994) AMA Erwin, 1994
Erwiniana eugeneae (Erwin, 1994) PUT Erwin, 1994
Erwiniana hilaris (Bates, 1871) AMA Martínez, 2003
Erwiniana notparkeri (Erwin, 1994)* PUT Erwin, 1994
Erwiniana sculpticollis (Bates, 1871) PUT Erwin, 1994
Trechinae Bonelli, 1810 Bembidiini Stephens, 1827 Erwiniana Paulsen & Smith, 2003 Erwiniana sulcicostis (Bates, 1882) CHO Martínez, 2003
Erwiniana wygo (Erwin, 1994)* CUN Erwin, 1994
Gouleta Erwin, 1994 Gouleta cayennensis (Dejean, 1831) ANT Erwin, 1973
Gouleta gentryi Erwin, 1994 AMA Martínez, 2003
Meotachys Erwin, 1974 Meotachys (Hylotachys) ballorum Boyd & Erwin, 2016 AMA Boyd & Erwin, 2016
Meotachys (Scolistichus) riparius Boyd & Erwin, 2016 AMA Boyd & Erwin, 2016
Pericompsus LeConte, 1852 Pericompsus (Eidocompsus) brasiliensis (Sahlberg, 1844) MAG Erwin, 1974
Pericompsus (Eidocompsus) immaculatus Bates, 1871 MAG Erwin, 1974
Pericompsus (Pericompsus) concinnus (Laferté-Sénectère, 1841) HUI Erwin, 1974
Pericompsus (Pericompsus) diabalius Erwin, 1974* MAG Erwin, 1974
Pericompsus (Pericompsus) gracilior (Bates, 1884) MAG Erwin, 1974
Pericompsus (Pericompsus) histrionellus Bates, 1884 MAG Erwin, 1974
Pericompsus (Pericompsus) reichei (Putzeys, 1845) MAG Erwin, 1974
Pericompsus (Pericompsus) silicis Erwin, 1974 CUN Erwin, 1974
Pericompsus metallicus Bates, 1871 HUI Erwin, 1974
Pogonini Laporte de Castelnau, 1834 Diplochaetus Chaudoir, 1872 Diplochaetus rutilus (Chevrolat, 1863) LAG Bousquet & Laplante, 1997; Reichardt, 1974a
Trechini Bonelli, 1810 Cnides Motschulsky, 1862 Cnides jeanneli Uéno, 1985 VAC Jeannel, 1958
Cnides rostratus Motschulsky, 1862 TOL Jeannel, 1958
Columbitrechus Mateu, 1982 Columbitrechus subsulcatus Mateu, 1982* CAU Mateu, 1982b
Oxytrechus Jeannel, 1927 Oxytrechus bousqueti Mateu, 1991* CAU Ruiz-Tapiador & Arenas, 2017
Oxytrechus campbelli Mateu, 1991* CAU Ruiz-Tapiador & Arenas, 2017
Oxytrechus caucaensis Mateu, 1991* CAU Ruiz-Tapiador & Arenas, 2017
Oxytrechus floresanus Giachino & Allegro, 2019* CAL Giachino et al., 2019
Oxytrechus jeanneli Mateu, 1991* CAU Ruiz-Tapiador & Arenas, 2017
Trechinae Bonelli, 1810 Trechini Bonelli, 1810 Oxytrechus Jeannel, 1927 Oxytrechus norae Mateu, 1982* CAU Ruiz-Tapiador & Arenas, 2017
Oxytrechus ruizianus Giachino & Allegro, 2019* CAL Giachino et al., 2019
Oxytrechus silvianus Mateu, 1991* CAU Ruiz-Tapiador & Arenas, 2017
Oxytrechus solitarius Mateu, 1991* CAU Ruiz-Tapiador & Arenas, 2017
Paratrechus Jeannel, 1920 Paratrechus incertus Mateu, 1999* CAU Mateu, 1998

Appendix 3 List of species of Cicindelidae Latreille, 1802 from Colombia until 2020, with their respective distribution within the country, species marked with asterisk (*) have been cited only from Colombia. Departments are codified according to ISO rule 3166-2. References are listed in appendix 1. 

Tribe Genus Species Department References
Cicindelini Latreille, 1802 Brasiella Rivalier, 1954 Brasiella (Brasiella) argentata (Fabricius, 1801) ANT, AMA, MET, PUT, VAC Cassola & Pearson 2001, Ramírez-Mora 2008, Arenas-Clavijo 2018
Brasiella (Brasiella) mendicula Rivalier, 1955 ANT, MET, NAR, SAN, VAC Cassola & Pearson 2001, Ramírez-Mora 2008, Arenas-Clavijo 2018
Brasiella (Brasiella) misella (Chaudoir, 1854) ANT, BOL, CES, CUN, MAG, SAN, NAR, VAC. Cassola & Pearson 2001, Ramírez-Mora 2008
Brasiella (Brasiella) nebulosa (Bates, 1874) VAC Cassola & Pearson 2001, Arenas-Clavijo 2018
Brasiella (Brasiella) venustula (Gory, 1833) ANT, COR, MET, NSA, SAN, VAC Cassola & Pearson 2001, Ramírez-Mora 2008, Arenas-Clavijo 2018
Callidema Guérin- Méneville, 1843 Callidema boussingaultii Guérin-Méneville, 1843 ANT, CAS, CAL, TOL, VAC Cassola & Pearson 2001, Ramírez-Mora 2008
Cenothyla Rivalier, 1969 Cenothyla consobrina (Lucas, 1857) AMA, MET Cassola & Pearson 2001
Cenothyla klichai Moravec, 2015 MET Moravec 2015
Cicindela Linnaeus, 1758 Cicindela (Cicindelidia) carthagena Dejean, 1831 BOL, CHO, VAC Cassola & Pearson 2001, Vitolo & Pearson 2003, Arenas-Clavijo 2018
Cicindela (Cicindelidia) favergeri Audouin & Brullé, 1839 CUN, MET, PUT, VAC Cassola & Pearson 2001, Arenas-Clavijo 2018
Cicindela (Cicindelidia) rufoaenea Horn, 1915 No data Cassola & Pearson 2001
Cicindela (Cicindelidia) trifasciata Fabricius, 1781 No data Cassola & Pearson 2001
Cylindera Westwood, 1831 Cylindera (Plectographa) suturalis ( Fabricius, 1798) No data Cassola & Pearson 2001
Habroscelimorpha Dokhtouroff, 1883 Habroscelimorpha auraria (Klug, 1834) LAG, MAG Rodriguez et al. 1994, Vítolo & Pearson 2003, Ramírez-Mora 2008
Habroscelimorpha schwarzi (Horn, 1923) VAC Cassola & Pearson 2001, Arenas-Clavijo 2018
Cicindelini Latreille, 1802 Langea Horn, 1901 Langea mellicollis Sumlin, 1993 AMA Vítolo & Pearson 2003
Mesacanthina Rivalier, 1969 Mesacanthina chalceola (Bates, 1872) MET Moravec 2020
Odontocheila Laporte de Castelnau, 1834 Odontocheila angulipenis Horn, 1932 MET, VID Cassola & Pearson 2001, Moravec 2013
Odontocheila batesii Chaudoir, 1860 AMA Vítolo & Pearson 2003, Arenas-Clavijo 2018
Odontocheila bipunctata (Fabricius, 1792) CAQ Moravec 2016, Vítolo 2004
Odontocheila cajennensis (Fabricius, 1787) No data Moravec 2016
Odontocheila chrysis ( Fabricius, 1801) ANT, RIS Vítolo & Pearson 2003
Odontocheila confusa (Dejean, 1825) ANT, AMA, MET, PUT Fernández et al 1993, Ramírez-Mora 2008
Odontocheila excisipenis Horn, 1932 ANT, CAU, CHO, NAR, VAC Cassola & Pearson 2001, Moravec 2012, Arenas-Clavijo 2018
Odontocheila eximia Lucas, 1857 AMA, PUT Vítolo & Pearson 2003
Odontocheila hamulipenis Horn, 1933* VAC Cassola & Pearson 2001, Moravec 2013, Arenas-Clavijo 2018
Odontocheila jordani Horn, 1898 CAU, NAR Cassola & Pearson 2001, Torres-Dominguez & Mendivil-Nieto 2012
Odontocheila margineguttata (Dejean, 1825) CAQ, GUA, GUV, MET Cassola & Pearson 2001
Odontocheila ochreata (Reiche, 1842) CAQ, MET, NAR, PUT Moravec 2016, Vítolo 2004
Odontocheila oseryi ( Lucas, 1857) AMA, PUT Moravec & Brzoska 2015, Vítolo 2004
Odontocheila salvini Bates, 1874 CHO, BOY, MAG Vítolo & Pearson 2003, Steinheil 1875b
Odontocheila simulator Horn, 1894* No data Wiesner 1992, Vítolo & Pearson 2003
Odontocheila trilbyana Thomson, 1857 AMA, VAU Vítolo & Pearson 2003
Opilidia Rivalier, 1954 Opilidia graphiptera ( Dejean, 1831) LAG, BOL, MAG Cassola & Pearson 2001, Ramírez-Mora 2008
Opilidia macrocnema (Chaudoir, 1852) CAU, NAR, VAC Cassola & Pearson 2001, Arenas-Clavijo 2018
Oxygonia Mannerheim, 1837 Oxygonia albitaenia Bates, 1871 No data Moravec 2015
Cicindelini Latreille, 1802 Oxygonia Mannerheim, 1837 Oxygonia fleutiauxi Horn, 1896 CAU Kippenhan 1997
Oxygonia floridula Bates, 1872 No data Vítolo & Pearson 2003
Oxygonia kippenhani Schüle, 2008* BOY Moravec 2017
Oxygonia kondratieffi Kippenhan, 1997 VAC Kippenhan 1997, Moravec 2017, Arenas-Clavijo 2018
Oxygonia moreti Deuve, 1992 NAR, RIS, VAC Kippenhan 1997, Moravec 2017, Arenas-Clavijo 2018
Oxygonia moronensis Bates, 1872 No data Cassola & Pearson 2001
Oxygonia nigricans Horn, 1926 CAU Kippenhan 1997, Moravec 2017
Oxygonia oberthueri Horn, 1896 ANT, NAR, RIS, VAC Cassola & Pearson 2001, Ramírez-Mora 2008, Arenas-Clavijo 2018
Oxygonia prodiga (Erichson, 1847) No data Kippenhan 1997, Moravec 2015
Oxygonia schoenherrii Mannerheim, 1837 ANT, CUN, VAC Kippenhan 1997, Moravec 2015
Oxygonia uniformis Horn, 1900 NAR Vítolo & Pearson 2003
Oxygonia vuillefroyi Chaudoir, 1869 BOY, CUN, HUI Vítolo & Pearson 2003
Pentacomia Bates, 1872 Pentacomia (Pentacomia) cupriventris ( Reiche, 1842) ANT, VAC Cassola & Pearson 2001, Ramírez-Mora 2008, Arenas-Clavijo 2018
Pentacomia ( Pentacomia) egregia (Chaudoir, 1835) AMA Cassola & Pearson 2001
Pentacomia (Poecilochila) lacordairei (Gory, 1833) AMA, GUV Cassola & Pearson 2001, Arenas-Clavijo 2018
Pentacomia (Poecilochila) ventralis (Dejean, 1825) ANT, CES, MET, VID Cassola & Pearson 2001, Ramírez-Mora 2008, Arenas-Clavijo 2018
Pentacomia chrysamma Bates, 1872 ANT Cassola & Pearson 2001
Ronhuberia Moravec & Kudrna, 2002 Ronhuberia fernandezi (Cassola, 2000) ANT Moravec & Kudrna 2002
Ctenostomatini Laporte de Castelnau, 1834 Ctenostoma Klug, 1821 Ctenostoma (Ctenostoma) succinctum (Laporte, 1834) GUV Vítolo & Pearson 2003
Ctenostomatini Laporte de Castelnau, 1834 Ctenostoma Klug, 1821 Ctenostoma (Neoprocephalus) maculicorne (Chevrolat, 1856) NAR, VAC Vítolo & Pearson 2003, Arenas-Clavijo 2018
Ctenostoma (Procephalus) dormeri Horn, 1898 ANT, VAC Naviaux 1998, Vítolo & Pearson 2003, Arenas-Clavijo 2018
Ctenostoma (Procephalus) ecuadorense Naviaux, 1998 CAU, VAC Naviaux 1998, Vítolo & Pearson 2003, Arenas-Clavijo 2018
Ctenostoma (Procephalus) longipalpe Naviaux, 1998 No data Naviaux 1998, Vítolo & Pearson 2003, Vítolo & Pearson 2003
Ctenostoma (Procephalus) maculosum Naviaux, 1998 CUN Naviaux 1998, Vítolo & Pearson 2003, Vítolo & Pearson 2003
Ctenostoma (Procephalus) nigrum Chaudoir, 1860 PUT, VAC Naviaux 1998, Vítolo & Pearson 2003, Vítolo & Pearson 2003
Ctenostoma (Procephalus) onorei Naviaux, 1998 CUN, VAC Naviaux 1998, Vítolo & Pearson 2003, Arenas-Clavijo 2018
Megacephalini Laporte de Castelnau, 1834 Aniara Hope, 1838 Aniara sepulchralis (Fabricius, 1801) CAQ, CAS, CUN, GUV, MET, TOL Cassola & Pearson 2001, Vítolo & Pearson 2003
Metriocheila Thomson, 1857 Metriocheila nigricollis (Reiche, 1842) HUI, PUT Cassola & Pearson 2001, Vítolo & Pearson 2003
Phaeoxantha Chaudoir, 1850 Phaeoxantha aequinoctialis (Dejean, 1825) CAS, MET Fernandez et al 1993, Vítolo & Pearson 2003
Phaeoxantha klugii Chaudoir, 1850 CAU, MET, PUT Fernandez et al 1993, Vítolo & Pearson 2003, Arenas-Clavijo 2018
Tetracha Hope, 1838 Tetracha (Neotetracha) affinis (Dejean, 1825) AMA, ANT, ATL, BOL, BOY, CAL, CHO, CUN, GUV, HUI, MAG, MET, SAN, TOL, VAC Wiesner 1992, Vítolo & Pearson 2003, Ramírez-Mora 2008
Tetracha (Neotetracha) cribrata Steinheil, 1875 ANT, BOL, CES, CUN, MET, NSA, SAN, TOL, VAC Steinheil 1875a, Vitolo & Pearson 2003, Arenas-Clavijo 2018
Tetracha (Neotetracha) fulgida (Klug, 1834) ANT, CAS, MET, PUT Cassola & Pearson 2001, Vítolo & Pearson 2003, Ramírez-Mora 2008
Tetracha (Neotetracha) gracilis (Reiche, 1842) No data Reiche 1842a
Tetracha (Neotetracha) lacordairei (Gory, 1833) ANT, BOY, CUN, HUI, MET, SAN Cassola & Pearson 2001, Ramírez-Mora 2008
Megacephalini Laporte de Castelnau, 1834 Tetracha Hope, 1838 Tetracha (Tetracha) carolina (Linnaeus, 1763) CAU, CES, RIS, VAC Cassola & Pearson 2001, Vítolo & Pearson 2003
Tetracha (Tetracha) sobrina (Dejean, 1831) AMA, ANT, BOL, BOY, CAL, CAQ, CAS, CAU, CHO, CUN, GUV, HUI, LAG, MET, QUI, RIS, SAN, TOL, VAC Vítolo & Pearson 2003, Ramírez-Mora 2008, Arenas-Clavijo 2018
Tetracha (Tetracha) spixii (Brullé, 1837) CHO, MET Fernández et al 1993, Vítolo & Pearson 2003
Tetracha (Tetracha) sommeri (Chaudoir, 1850) No data Chaudoir 1850
Oxycheilini Chaudoir, 1860 Cheiloxya Guérin- Méneville, 1855 Cheiloxya binotata (Laporte, 1833) No data Fernandez et al 1993, Vítolo & Pearson 2003
Oxycheila Dejean, 1825 Oxycheila aquatica Guérin-Méneville, 1843* CUN, SAN, VAC Wiesner 1999
Oxycheila binotata Gray, 1832 No data Wiesner 1999
Oxycheila brzoskai Wiesner, 1999* ANT, NAR, VAC Cassola & Pearson 2001, Vítolo & Pearson 2003, Ramírez-Mora 2008
Oxycheila chestertonii Bates, 1872 ANT, CUN, MET, TOL, VAC Wiesner 1999, Vítolo & Pearson 2003, Ramírez-Mora 2008
Oxycheila femoralis Laporte, 1833 CUN, TOL Vítolo & Pearson 2003
Oxycheila gracillima Bates, 1872 AMA Vítolo & Pearson 2003
Oxycheila haenschi Horn, 1900 AMA Vítolo & Pearson 2003
Oxycheila howdeni Brouerius van Nidek, 1980* ANT, BOL, CAU, VAC Wiesner 1999, Vítolo & Pearson 2003, Arenas-Clavijo 2018
Oxycheila pearsoni Wiesner, 1999* ANT, CAU Wiesner 1999, Vítolo & Pearson 2003
Oxycheila polita Bates, 1872 No data Wiesner 1999, Vítolo & Pearson 2003
Oxycheila pseudoaquatica Wiesner, 1999* VAC Wiesner 1999, Vítolo & Pearson 2003
Oxycheila pseudostrandi Wiesner, 1999* CAU, NAR Vítolo & Pearson 2003
Oxycheila tristis (Fabricius, 1775) No data Wiesner 1999
Pseudoxycheila Guérin-Méneville, 1839 Pseudoxycheila atahualpa Cassola, 1997 BOY, PUT, SAN Cassola 1997
Oxycheilini Chaudoir, 1860 Pseudoxycheila Guérin-Méneville, 1839 Pseudoxycheila bipustulata (Latreille, 1811)* ANT, BOY, CAL, CAU, CES, CHO, COR, CUN, HUI, MAG, MET, NAR, NSA, PUT, QUI, RIS, SAN, SUC, TOL, VAC Vítolo & Pearson 2003, Ramírez-Mora 2008, Arenas-Clavijo 2018
Pseudoxycheila caribe Cassola, 1997 NSA Cassola 1997
Pseudoxycheila ceratoma Chaudoir, 1865* No data Vítolo & Pearson 2003
Pseudoxycheila chaudoiri Dokhtouroff, 1882* BOY, CAL, CAU, MET, QUI, RIS, VAC Cassola 1997, Vítolo & Pearson 2003
Pseudoxycheila columbiana Cassola, 1997* BOY, CAU, CUN, MAG, MET, SAN, TOL, VAC Cassola 1997, Vítolo & Pearson 2003
Pseudoxycheila confusa Cassola, 1997 ANT, BOY, CAU, CES, CAL, CAS, CHO, CAQ, CUN, HUI, MET, RIS, SAN, TOL, VAC Cassola 1997, Vítolo & Pearson 2003
Pseudoxycheila lateguttata Chaudoir, 1844 ANT, BOY, CAU, HUI Cassola 1997, Vítolo & Pearson 2003
Pseudoxycheila macrocephala Cassola, 1997* ANT, CAL, PUT Cassola 1997, Vítolo & Pearson 2003
Pseudoxycheila nitidicollis Cassola, 1997 HUI, PUT Cassola 1997, Vítolo & Pearson 2003
Pseudoxycheila tarsalis Bates, 1869* No data Cassola & Pearson 2001, Vítolo & Pearson 2003

Received: April 07, 2021; Accepted: June 28, 2021

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