0123-3475
0123-3475
S0123-34752021000200047
10.15446/rev.colomb.biote.v23n2.88487
Costa Rica
Costa Rica
00
12
2021
00
12
2021
23
2
47
66
[
2008
27
4
4
687-98
]
[
2013
14
4
4
7815-28
]
[
2011
60
2
2
403-14
]
[
2010
28
1
1
169-83
]
[
2014
8
3
3
295-304
]
[
2018
9
9
801
]
[
2017
254
1
1
3-16
]
[
2011
77
4
4
607-46
]
[
2019
14
5
5
]
[
2008
27
3
3
411-24
]
[
2014
8
3
3
413-21
]
[
2012
159
1
1
266-85
]
[
2002
32
4
4
715-23
]
[
2014
26
62-70
]
[
2016
17
1
1
312
]
[
2010
229-50
]
[
2014
233-52
]
[
2016
76
76
105-24
]
[
2015
56
5
5
917-29
]
[
2005
169
1
1
75-82
]
[
2015
206
3
3
775-83
]
[
2015
84
1
1
56-69
]
[
2009
36
1
1
17-29
]
[
2011
10
68
68
15253-8
]
[
2019
10
2
2
142
]
[
2000
5
5
5
199-206
]
[
2015
72
4
4
673-89
]
[
FAO
2016
]
[
2012
1-33
]
[
2010
15
7
7
409-17
]
[
2000
12
12
901-015
]
[
2020
11
11
139
]
[
2013
31
7
7
397-405
]
[
2009
28
2
2
301-11
]
[
2020
168
1
1
98-117
]
[
2011
168
11
11
1286-94
]
[
2016
6
1
1
24978
]
[
2017
8
8
1657
]
[
2011
30
3
3
297-310
]
[
2013
31
3
3
559-73
]
[
2005
169
4
4
746-52
]
[
2014
65
1
1
715-41
]
[
2011
27
2
2
297-306
]
[
1994
6
1277-87
]
[
2002
7
3
3
106-11
]
[
2002
21
2
2
141-6
]
[
2017
59
2
2
86-101
]
[
2017
118
118
45-54
]
[
2019
1
1-24
]
[
2008
146
2
2
623-35
]
[
2018
41
5
5
898-907
]
[
2010
103
2
2
267-77
]
[
2016
283
1832
1832
20160792
]
[
2018
9
9
346
]
[
2012
12
1
1
74
]
[
2011
6
8
8
]
[
2018
333-42
]
[
2015
126
126
23-34
]
[
1967
66
2
2
143-84
]
[
2018
20
4
4
464-73
]
[
2014
32
1
1
42-51
]
[
2019
19
1
1
38
]
[
2016
6
1
1
35040
]
[
2017
89
3
3
510-26
]
[
2019
10
10
772
]
[
2007
20
3
3
233-48
]
[
2013
112
2
2
257-62
]
[
2018
12
6
6
399-408
]
[
2012
63
2
2
599-616
]
[
2015
6
1
1
8326
]
[
2015
175
1
1
1-8
]
[
2000
46
4
4
873-83
]
[
2007
130
2
2
218-29
]
[
2015
89
4-5
4-5
421-31
]
[
2018
98
1
1
137-52
]
[
2012
1819
2
2
97-103
]
[
2014
9
5
5
]
[
2015
33
8
8
862-9
]
[
2013
162
3
3
1720-32
]
[
2016
6
1
1
26685
]
[
2017
7
4
4
239
]
[
2016
38
7
7
181
]
[
2017
8
8
1881
]
[
2019
1
25-52
]
[
2010
467
7311
7311
43-51
]
[
2011
52
S4
S4
S453-70
]
[
2004
a
166
1
1
141-9
]
[
2004
b
2
6
6
477-86
]
[
2011
33
4
4
1521-32
]
[
2016
38
12
12
1991-2006
]
[
2001
28
2
2
123-33
]
[
2011
14
3
3
232-9
]
[
1995
29
29
691-702
]
[
2002
25
2
2
163-71
]
[
2014
32
4
4
347-55
]
[
2014
9
12
12
]
[
2016
7
7
935
]
[
2017
355
6330
6330
1122-3
]
[
2012
31
9
9
1659-65
]
[
2015
120
3
3
1139-57
]
[
2007
176
1
1
70-81
]
[
2019
11
1
1
35
]
[
2017
15
2
2
207-16
]
[
2015
169
1
1
266-82
]
[
2019
13
4
4
409-18
]
[
2011
155
1
1
421-32
]
[
2016
7
7
817
]
[
2015
6
6
895
]
[
2015
14
3
3
407-26
]
[
2018
274
274
369-82
]
[
2015
58
4
4
568-74
]
[
2008
227
2
2
299-308
]
[
2001
28
4
4
397-408
]
[
2003
15
9
9
2076-92
]
[
2015
58
1
1
52-60
]
[
2017
28
1
1
15-28
]
[
2018
16
2
2
354-66
]
[
2015
6
6
84
]
[
2007
2
3
3
135-8
]
[
1975
]
[
2020
47
3
3
1573-81
]
[
2011
16
7
7
363-71
]
[
2016
28
3
3
786-803
]
[
2012
]
[
2017
a
246
3
3
453-69
]
[
2011
]
[
2017
b
65
39
39
8674-82
]
[
2003
218
1
1
1-14
]
[
2016
4
3
3
162-76
]
[
2017
257
257
74-83
]
[
2014
36
7
7
1841-51
]
[
2017
51
5
5
371-83
]
[
1997
16
17
17
5247-59
]
[
2019
249
6
6
1963-75
]
[
2016
14
2
2
592-602
]
[
2011
a
234
3
3
541-54
]
[
2020
11
11
14
]
[
2009
23
2
2
289-98
]
[
2011
b
181
1
1
1-16
]
[
2015
236
236
214-22
]
[
2010
26
3
3
455-65
]
[
2016
34
1
1
353-64
]